Pedicularis attollens |
Pedicularis groenlandica |
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attol lousewort, attoll lousewort, elephant snouts, little elephant head, little elephant's head, woolly Mammoth |
bull elephant's-head, elephant's head, elephant's head pedicularis, elephant's-head lousewort, elephant-head lousewort, pédiculaire du groenland |
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Habit | Plants 15–78 cm. | Plants 10–60 cm. | ||||
Leaves | basal 5–25, blade elliptic, 60–150(or 200–250) x 3–23 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous or scattered glands; cauline 2–20, blade elliptic, 5–50(–100) x 1–5 mm, undivided or 1(or 2)-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous. |
basal 5–20, blade lanceolate, 20–150 x 5–25 mm, 1-pinnatifid or slightly 2-pinnatifid, margins of adjacent lobes nonoverlapping, 1- or 2-serrate, surfaces glabrous; cauline 3–31, blade lanceolate, 10–150 x 1–25 mm, 1-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous. |
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Racemes | simple, 1–3, exceeding basal leaves, each 10–50-flowered; bracts lanceolate to triangular, 5–10 x 3–10 mm, pinnatifid, margins entire, surfaces glabrous or tomentose. |
simple, 1 or 2, exceeding basal leaves, each 20–75-flowered; bracts linear to trullate, 5–10 x 2–10 mm, undivided to pinnatifid, margins entire, serrate, or 2-serrate, surfaces glabrous. |
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Pedicels | 1.2–1.6 mm. |
0.5–1 mm. |
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Flowers | calyx 4–5 mm, glabrous or tomentose, lobes 5, triangular, 2–2.5 mm, apex entire, glabrous; corolla 6–8 mm, tube pink, rarely white, 3–6 mm; galea white or pink with 2 purple spots or stripes, 1–2 mm, beaked, beak coiled, 3–6 mm, base curving, margins entire medially and distally, apex not surrounded by abaxial lip, axis of coil nearly vertical; abaxial lip pendulous, white or pink with purple stripe, 4–5.5 mm. |
calyx 3–5 mm, glabrous or hispid, lobes 5, deltate, 0.5–1.5 mm, apex entire, glabrous; corolla 5–8 mm, tube purple, rarely white, 3–5 mm; galea pink to purple, rarely white, 1.5–3 mm, beaked, beak coiled, 5–18 mm, base curving, margins entire medially and distally, apex not surrounded by abaxial lip, axis of coil nearly horizontal; abaxial lip pendulous, purple, rarely white, 2–5 mm. |
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2n | = 16. |
= 16. |
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Pedicularis attollens |
Pedicularis groenlandica |
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Phenology | Flowering Jun–Sep. | |||||
Habitat | Montane and alpine to arctic bogs, fens, marshes, and forested swamps, seepage areas, stream banks, fens, clay gravel flood plains of rivers. | |||||
Elevation | 600–3500 m. (2000–11500 ft.) | |||||
Distribution |
CA; NV; OR
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AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; LB; MB; NU; ON; QC; SK; YT; Greenland
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Discussion | Subspecies 2 (2 in the flora). The flowers of Pedicularis attollens, like those of P. groenlandica, resemble an elephant’s head, and A. Heller placed them both in Elephantella. The short, upturned beak, in contrast to the long, more horizontal downturned beak of P. groenlandica, is a distinguishing feature of P. attollens. Whereas P. groenlandica occurs across much of western and arctic North America, P. attollens is found primarily in the Cascade Range of central and southern Oregon and the Sierra Nevada of California and Nevada. It is also reported from the Klamath Range to the west and the White and Sweetwater mountains and the Warner Range to the east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pedicularis groenlandica has a domed galea, a long, curved beak, and relatively large lateral lobes of the abaxial lip that remarkably resemble the head, trunk, and ears of an elephant. Pedicularis groenlandica is the most widely distributed Pedicularis species in North America; it occurs from the southern Sierra Nevada and southern Rocky Mountains well into the Arctic and Greenland. Pedicularis attollens and P. groenlandica are sympatric in the Sierra Nevada and the Cascades of central and southern Oregon and can be difficult to distinguish. Pedicularis groenlandica is distinguished by the size and orientation of the beak and color of the corolla; the beak is over twice the length of the beak of P. attollens and is oriented more horizontally compared to the upturned beak of P. attollens. Beak orientation is very difficult to ascertain in herbarium material due to flattening; beak length is a better character on herbarium specimens. The corollas of P. groenlandica are purple with lighter purple to pinkish abaxial lips, whereas the corollas of P. attollens vary from pink to light pink often with conspicuous dark purple spots on the galeas. White-flowered forms of P. groenlandica (forma pallida Lepage) are occasionally seen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 514. | FNA vol. 17, p. 523. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Elephantella attollens | Elephantella groenlandica, P. groenlandica subsp. surrecta | ||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 384. (1868) | Retzius: Fl. Scand. Prodr. ed. 2, 145. (1795) | ||||
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