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attol lousewort, attoll lousewort, elephant snouts, little elephant head, little elephant's head, woolly Mammoth

Chamisso's lousewort

Habit Plants 15–78 cm. Plants 10–25 cm.
Leaves

basal 5–25, blade elliptic, 60–150(or 200–250) x 3–23 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous or scattered glands;

cauline 2–20, blade elliptic, 5–50(–100) x 1–5 mm, undivided or 1(or 2)-pinnatifid, margins of adjacent lobes nonoverlapping, serrate, surfaces glabrous.

basal 5–15, blade lanceolate, 15–40 x 10–20 mm, 2-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, serrate, surfaces glabrous;

cauline whorled, whorls 1–4, leaves per whorl 3–5, blade lanceolate, 10–60 x 10–30 mm, 1- or 2-pinnatifid, margins of adjacent lobes nonoverlapping or slightly overlapping distally, entire or serrate, surfaces glabrous.

Racemes

simple, 1–3, exceeding basal leaves, each 10–50-flowered;

bracts lanceolate to triangular, 5–10 x 3–10 mm, pinnatifid, margins entire, surfaces glabrous or tomentose.

verticillate, 1–3, exceeding basal leaves, each 8–20-flowered;

bracts lanceolate to subulate, 10–30 x 5–10 mm, 1-pinnatifid or undivided, margins serrate to 2-serrate, surfaces glabrous.

Pedicels

1.2–1.6 mm.

1.5–2.5 mm.

Flowers

calyx 4–5 mm, glabrous or tomentose, lobes 5, triangular, 2–2.5 mm, apex entire, glabrous;

corolla 6–8 mm, tube pink, rarely white, 3–6 mm;

galea white or pink with 2 purple spots or stripes, 1–2 mm, beaked, beak coiled, 3–6 mm, base curving, margins entire medially and distally, apex not surrounded by abaxial lip, axis of coil nearly vertical;

abaxial lip pendulous, white or pink with purple stripe, 4–5.5 mm.

calyx 6–8 mm, glabrous, lobes 5, triangular, 0.5–1.4 mm, apex entire, ciliate;

corolla 18–20 mm, tube white to pink, 12–15 mm;

galea purple, 6–8 mm, beaked, beak straight, 1–3 mm, margins entire medially and distally, apex extending beyond abaxial lip;

abaxial lip pink or purple, 4–5 mm.

2n

= 16.

Pedicularis attollens

Pedicularis chamissonis

Phenology Flowering Jul–Sep.
Habitat Arctic meadows, tundras.
Elevation 10–300 m. (0–1000 ft.)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; e Asia
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

The flowers of Pedicularis attollens, like those of P. groenlandica, resemble an elephant’s head, and A. Heller placed them both in Elephantella. The short, upturned beak, in contrast to the long, more horizontal downturned beak of P. groenlandica, is a distinguishing feature of P. attollens. Whereas P. groenlandica occurs across much of western and arctic North America, P. attollens is found primarily in the Cascade Range of central and southern Oregon and the Sierra Nevada of California and Nevada. It is also reported from the Klamath Range to the west and the White and Sweetwater mountains and the Warner Range to the east of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pedicularis chamissonis differs from P. verticillata by having leaves and flowers about 1.5 times larger with a galea that is extended into a conical beak, leaves lanceolate versus elliptic, and lateral lobes of the abaxial lip more than twice the size and more pendulous.

Pedicularis chamissonis is chiefly an Asian species, found on the Kamchatka Peninsula, Kuril Islands, Sakhalin Island, and Japan; the only North American populations occur in the Aleutian Islands of Alaska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves 1-pinnatifid.
subsp. attollens
1. Basal leaves 2-pinnatifid.
subsp. protogyna
Source FNA vol. 17, p. 514. FNA vol. 17, p. 519.
Parent taxa Orobanchaceae > Pedicularis Orobanchaceae > Pedicularis
Sibling taxa
P. angustifolia, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. chamissonis, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
P. angustifolia, P. attollens, P. aurantiaca, P. bracteosa, P. canadensis, P. capitata, P. centranthera, P. contorta, P. crenulata, P. cystopteridifolia, P. densiflora, P. dudleyi, P. flammea, P. furbishiae, P. groenlandica, P. hirsuta, P. howellii, P. labradorica, P. lanata, P. lanceolata, P. langsdorffii, P. lapponica, P. oederi, P. ornithorhyncha, P. ornithorhynchos, P. palustris, P. parryi, P. parviflora, P. pennellii, P. procera, P. pulchella, P. racemosa, P. rainierensis, P. semibarbata, P. sudetica, P. sylvatica, P. verticillata
Subordinate taxa
P. attollens subsp. attollens, P. attollens subsp. protogyna
Synonyms Elephantella attollens
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 384. (1868) Steven: Mém. Soc. Imp. Naturalistes Moscou 6: 20, plate 4, fig. 1. (1822)
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