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cross-shape mitrewort, side-flower mitrewort, smallflower miterwort

creeping mitrewort, slightstem miterwort

Habit Plants not stoloniferous. Plants usually stoloniferous.
Flowering stems

10–55 cm.

13–45 cm.

Leaves

petiole 2–15 cm, short stipitate-glandular and long stipitate-glandular, longer hairs retrorse or, sometimes, spreading, white, tan, or brown;

blade cordate or reniform, shorter than or sometimes ± as long as wide, (1.5–)2.5–10 × 2–8(–12) cm, margins 5- or 7-lobed, singly or doubly crenate, uniformly ciliate, apex of terminal lobe rounded to obtuse, surfaces glabrous or short stipitate-glandular and sparsely long stipitate-glandular;

cauline leaves absent.

petiole 1.5–14 cm, short stipitate-glandular and, usually, long stipitate-glandular, longer hairs spreading to retrorse, white;

blade broadly cordate to broadly ovate, ± as long as wide, 1.2–6.5(–8.5) × 1.4–7(–8.2) cm, margins 3-, 5-, or 7-lobed, crenate-dentate, irregularly ciliate, apex of terminal lobe acute to obtuse, surfaces short stipitate-glandular and sparsely long stipitate-glandular;

cauline leaves 1–3, proximal or mid cauline, alternate, sessile or petiolate, blade 0.3–4.3(–6) × 0.4–4.4(–7) cm.

Inflorescences

1–3(–6), closely 10–45-flowered, 1 flower per node, strongly secund, 10–55 cm, short stipitate-glandular and sparsely long stipitate-glandular proximally and distally.

1–2(–4), remotely 5–25-flowered, 1(–2) flowers per node, (anthesis basipetalous), not secund, 13–45 cm, short stipitate-glandular and sparsely long stipitate-glandular proximally, short stipitate-glandular distally.

Pedicels

0.5–1.5 mm, short stipitate-glandular.

2–8 mm, short stipitate-glandular.

Flowers

hypanthium turbinate-campanulate, 1.4–2.7 × 1.5–2.5(–3) mm;

sepals erect or with tips spreading, greenish white to white, sometimes purple-tinged, oblong, oblong-obovate, or widely ovate, 1–2.2 × 0.7–1.2 mm;

petals white, sometimes purple-tinged, 3-lobed or, sometimes, unlobed, 1.5–4 mm, lobes linear to lanceolate, lateral lobes ascending or spreading;

stamens 5, opposite sepals;

filaments white, 0.1–0.2 mm;

anthers 0.3–0.5 × 0.2–0.3 mm;

ovary ca. 2/3–3/4 inferior;

styles divergent, flattened, 0.1–0.2 mm;

stigmas unlobed.

hypanthium broadly campanulate, 1.5–2 × 3–5.5 mm;

sepals spreading or recurved, greenish, deltate-ovate, 0.9–1.5 × 1.4–2 mm;

petals whitish green or greenish, sometimes purple tinged proximally, 5–9-lobed, 3–4 mm, lobes linear, lateral lobes spreading;

stamens 5, opposite sepals;

filaments usually purple proximally and white distally, sometimes entirely white, 0.6–0.9 mm;

anthers 0.2–0.4 × 0.2–0.4 mm;

ovary 1/2 inferior;

styles erect or spreading, cylindric, 0.8–1.1 mm;

stigmas unlobed.

Seeds

dark reddish brown to blackish, 0.8–1.2 mm, pitted.

reddish purple or dark reddish brown, 0.7–1 mm, pitted.

2n

= 14.

= 14.

Mitella stauropetala

Mitella caulescens

Phenology Flowering May–Jul. Flowering Apr–Jul.
Habitat Open to dense woods, seasonally moist, springy sites Deep woods, meadows, stream banks
Elevation 700-3000 m (2300-9800 ft) 0-2100 m (0-6900 ft)
Distribution
from FNA
CO; ID; MT; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; BC
[WildflowerSearch map]
Discussion

Two varieties of Mitella stauropetala have been recognized. Plants from Oregon and Washington have been referred to var. stauropetala, characterized by hypanthium plus sepals often over 3 mm and petal blades with linear lobes. In northern Colorado, southeastern Idaho, eastern Utah, and Wyoming, var. stauropetala is replaced by var. stenopetala, with hypanthium plus sepals rarely over 3 mm and petal blades less deeply trifid (sometimes entire) and with broader lateral lobes. Variety stenopetala is morphologically similar in many respects to M. trifida. Mitella trifida and M. stauropetala require study to determine if plants referred to var. stenopetala are the result of hybridization or integradation between the two species.

D. E. Soltis and P. S. Soltis (1986) found morphologic and allozymic evidence of fertile intergeneric hybrids between Mitella stauropetala and Conimitella williamsii in Wyoming. Data from cpDNA analysis suggest that C. williamsii is derived from within the M. diversifoliaM. stauropetalaM. trifida alliance (D. E. Soltis et al. 1990).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 113. FNA vol. 8, p. 112.
Parent taxa Saxifragaceae > Mitella Saxifragaceae > Mitella
Sibling taxa
M. breweri, M. caulescens, M. diphylla, M. diversifolia, M. nuda, M. ovalis, M. pentandra, M. trifida
M. breweri, M. diphylla, M. diversifolia, M. nuda, M. ovalis, M. pentandra, M. stauropetala, M. trifida
Synonyms M. stauropetala var. stenopetala, Ozomelis stauropetala Mitellastra caulescens
Name authority Piper: Erythea 7: 161. 1899 , Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 586. 1840 ,
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