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cotton-seed, falzblume, micrope

cottontop, q-tips, slender cotton-weed, slender cottonseed

Habit Annuals, 1–50 cm. Plants 1–50 cm.
Stems

1, ± erect, or 2–5[–10], ascending to erect [prostrate].

Leaves

cauline; mostly alternate [opposite];

blades narrowly oblanceolate to elliptic [spatulate].

Involucres

inconspicuous.

Receptacles

depressed-spheric or obovoid (heights 0.5–1.8 times diams.), glabrous.

depressed-spheric, mostly 0.3–0.6 mm, heights 0.5–0.8 times diams.

Pistillate florets

4–12.

Bisexual florets

0.

Phyllaries

4–6, ± equal (unlike paleae, scarious, hyaline).

Heads

usually in glomerules of 2–5 in racemiform to paniculiform or distally ± dichasiform [axillary] arrays, sometimes borne singly.

depressed-spheric, 2–4 × 3–6 mm.

Cypselae

brown, monomorphic: ± trigonously [evenly] compressed, ± obovoid, curved, gibbous abaxially, faces glabrous, smooth, shiny, corolla scars ± lateral;

pappi: pistillate 0, staminate 0 or of 1–5 bristles (hidden in heads).

1.4–2.6 mm, corolla scars ± median;

pappi: staminate 0 or of 1, ± smooth bristle 0.9–1.5 mm.

Pistillate

paleae falling, erect to incurved;

bodies with 5+ nerves (nerves ± parallel, obscure), obovoid, saccate most of lengths (trigonously [evenly] compressed, galeate, abaxially rounded [corniculate-crested], each enclosing a floret);

wings ± erect (and lateral) or inflexed (and subapical).

paleae 4–7(–8) in 1 series, longest 2–4 mm;

wings obscure, withering, lateral, ± erect, narrowly oblanceolate, involute;

bodies galeate, cartilaginous to bony throughout, sericeous to lanuginose.

Staminate

paleae 0 or 1–3, falling, erect in fruit (not enlarged), shorter than pistillate paleae;

bodies linear-lanceolate to oblanceolate.

paleae 0.;

staminate corollas 1–2 mm, lobes usually 5.

Functionally

staminate florets 2–5;

corolla lobes 4–5, ± equal.

x

= 14.

Micropus

Micropus californicus

Distribution
from USDA
w United States; nw Mexico; s Europe; sw Asia; n Africa
[BONAP county map]
from FNA
CA; OR; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 5 (2 in the flora).

See discussion of Filagininae following the tribal description (p. 385).

Micropus species are found mostly in dry, open habitats of Mediterranean climates. In the flora, they are known only from west-draining portions of the Californian Floristic Province and the Willamette Valley in Oregon.

The two North American species constitute Micropus sect. Rhyncholepis Nuttall. Recent European workers (e.g., J. Holub 1998) have included sect. Rhyncholepis in Bombycilaena, leaving M. supinus Linnaeus in a monotypic genus. Based on phylogenetic data (J. D. Morefield 1992), that approach would include in Bombycilaena species ancestral to, and derived from ancestors of, Micropus. I maintain Micropus in its traditional sense here. Micropus and Psilocarphus appear to be monophyletic sister genera derived from near or within Stylocline. A malformed specimen from Monterey County, California, appears to be a sterile hybrid between M. californicus and a species of Psilocarphus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Though common and often found on recent disturbances within its range, Micropus californicus does not appear to be particularly invasive or weedy. The mature pistillate paleae, with roughly the profile of a harp or of an inverted lower-case letter “q,” are distinctive. That shape and the accompanying dense, cottony indument of the common variety explain the vernacular names. By contrast, pistillate paleae of M. amphibolus have roughly the profile of a human head wearing a billed cap.

Within the more limited range of var. subvestitus, the two varieties are broadly sympatric, usually in separate populations, occasionally in mixed populations that sometimes include intermediate plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pistillate paleae 8–12 in 2 series, bodies mainly chartaceous, cartilaginous medially, wings prominent, subapical, inflexed, plane to concave; cypselae: corolla scars in distal 1/4; receptacle heights 1.2–1.8 times diams.; staminate paleae mostly 1–3; staminate corolla lobes 4 (–5); staminate pappi of 1–5 bristles
M. amphibolus
1. Pistillate paleae 4–7(–8) in 1 series, bodies cartilaginous to bony throughout, wings obscure, lateral, ± erect, involute; cypselae: corolla scars ± median; receptacle heights 0.5–0.8 times diams.; staminate paleae 0; staminate corolla lobes usually 5; staminate pappi 0 or of 1 bristle
M. californicus
1. Pistillate paleae densely and loosely lanuginose, longest usually 3–4 mm
var. californicus
1. Pistillate paleae thinly sericeous-lanuginose, longest usually 2–3 mm
var. subvestitus
Source FNA vol. 19, p. 454. Author: James D. Morefield. FNA vol. 19, p. 455.
Parent taxa Asteraceae > tribe Gnaphalieae Asteraceae > tribe Gnaphalieae > Micropus
Sibling taxa
M. amphibolus
Subordinate taxa
M. amphibolus, M. californicus
M. californicus var. californicus, M. californicus var. subvestitus
Synonyms Bombycilaena Bombycilaena californica
Name authority Linnaeus: Sp. Pl. 2: 927. (1753): Gen. Pl. ed. 5, 398. (1754) Fischer & C. A. Meyer: Index Seminum (St. Petersburg) 2: 42. (1836)
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