Lilium pardalinum |
Lilium parryi |
|||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
California tiger lily, leopard lily, panther lily |
lemon lily, Parry lily |
|||||||||||||||||
Bulbs | rhizomatous, usually branching, continuously scaly, 1.4–5.1 × 3.9–19 cm, 0.2–0.6 times taller than long; scales sometimes unsegmented but always some 2–4-segmented on each bulb, longest 1–3.3 cm; stem roots absent. |
rhizomatous, unbranched, continuously scaly, 1.5–4.7 × 3.5–11 cm, 0.2–0.6 times taller than long; scales (1–)2(–4)-segmented, longest 0.9–3.7 cm; stem roots absent. |
||||||||||||||||
Stems | to 2.8 m, strongly clonal and thus forming dense colonies, to weakly clonal and forming small colonies or clumps. |
to 1.9 m. Buds rounded in cross section. |
||||||||||||||||
Buds | rounded in cross section. |
|||||||||||||||||
Leaves | usually ± evenly distributed along stem, rarely concentrated proximally, scattered or in 1–6 whorls or partial whorls, 3–19 leaves per whorl, horizontal and drooping at tips to ascending, 4.9–26.5 × 0.3–5.6 cm, 3–34 times longer than wide; blade usually ± elliptic, wide or narrow, margins usually not undulate, apex acute, often narrowly so; veins and margins ± smooth abaxially. |
occasionally scattered in young plants or in 1–5 whorls or partial whorls, 3–18 leaves per whorl, ± horizontal and drooping at tips or ascending, 7.8–29 × 0.5–4.9 cm, 2.6–29 times longer than wide; blade elliptic to narrowly linear, or ± obovate, often lanceolate in distal leaves, margins not undulate, apex acute, often narrowly so; veins and margins ± smooth abaxially. |
||||||||||||||||
Inflorescences | racemose, 1–28(–35)-flowered. |
racemose, 1–31-flowered. |
||||||||||||||||
Flowers | pendent, usually not fragrant; perianth Turk’s-cap-shaped; sepals and petals reflexed 1/4–1/3 along length from base, yellow, yellow-orange, or orange proximally, darker orange to red-orange to red on distal 1/5–3/5 (entirely orange or yellow-orange in subsp. wigginsii), with maroon spots concentrated proximally and always surrounded by yellow or orange if extending into distal reddish zone, conspicuously green abaxially on proximal ± 1/5, not distinctly clawed; sepals not ridged abaxially, 3.5–10.4 × 0.9–2.2 cm; petals 3.4–10.2 × 0.9–2.5 cm; stamens moderately to strongly exserted; filaments moderately to widely spreading, diverging 7°–22° from axis; anthers ± magenta or sometimes orange, orange-pink, or pale yellow, 0.5–2.2 cm; pollen red-brown, red-orange, brown-orange, rust, orange, or yellow; pistil 3.1–7.5 cm; ovary 1–2.2 cm; style green, often pale, rarely sordid; pedicel 6–32 cm. |
opening before dusk, horizontal or somewhat nodding, slightly bilaterally symmetric, strongly fragrant; perianth funnelform; sepals and petals recurved 3/5 along length from base, lower less recurved than upper and forming landing platform, bright yellow with sparse, usually minute maroon spots, not distinctly clawed; sepals not ridged abaxially, oblanceolate, 7.7–10.7 × 1.1–1.7 cm; petals noticeably wider than sepals, often very wide distally, 7.8–10.6 × 1.1–2.1 cm, apex widely acute or sometimes obtuse; stamens barely exserted; filaments barely spreading, diverging at 5°–12°; anthers pale magenta-brown, 0.8–1.4 cm; pollen rust-orange or orange-brown; pistil 5.3–9.3 cm; ovary 1.6–2.9 cm; style green, often pale; pedicel 2–17.5 cm. |
||||||||||||||||
Capsules | 2.2–5.7 × 1.2–2.1 cm, 1.5–3.7 times longer than wide. |
3.9–5.9 × 1.1–1.7 cm, 2.5–4.4 times longer than wide. |
||||||||||||||||
Seeds | 123–264. |
141–303. |
||||||||||||||||
2n | = 24. |
|||||||||||||||||
Lilium pardalinum |
Lilium parryi |
|||||||||||||||||
Phenology | Flowering summer (late May–early Sep). | |||||||||||||||||
Habitat | Meadows, streams, and willow (Salix spp.) thickets in mixed conifer forests | |||||||||||||||||
Elevation | 1300–2600 m (4300–8500 ft) | |||||||||||||||||
Distribution |
CA; OR
|
AZ; CA
|
||||||||||||||||
Discussion | Subspecies 5 (5 in the flora). The subspecies of Lilium pardalinum display a classic pattern of discrete geographical ranges with intervening zones of introgression, and no two occur sympatrically without intermixing. Plants in the hybrid zones are confusing in appearance and cannot be assigned to subspecies. However, each subspecies is fairly well marked within its core distribution. With the exception of subsp. pitkinense, the subspecies of L. pardalinum can be common plants in the proper habitats within their rather narrow distributions. Leaf size and shape are quite variable in Lilium pardalinum subspecies and often clearly dependent on environment. In populations that typically have narrow, ascending leaves, shaded plants often have wide, horizontal leaves. This hampers taxonomic separation as well as identification, especially of herbarium specimens. Further field study is desirable. Lilium pardalinum is primarily pollinated by western tiger swallowtails (Papilio rutulus Lucas, family Papilionidae) and pale swallowtails (P. eurymedon Lucas); several species of hummingbirds (family Trochilidae) are also important visitors, especially when butterflies are rare. The Atsugewi, Karok, and Yana ate Lilium pardalinum bulbs steamed or baked in an earth oven (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants in Arizona start to bloom in May (T. H. Kearney and R. H. Peebles 1960), while the majority of California plants flower in July and August. Plants from the San Gabriel Mountains of California sometimes have wider leaves and have been given status as Lilium parryi var. kessleri, but this variation is due primarily to the rather shaded habitat of many of these populations. No significant vegetative discontinuity can be recognized across the range of this species, so no varieties are recognized here. Lilium parryi probably arose from an ancestor in common with L. pardalinum (M. W. Skinner 1988), and subsequently diverged to become pollinated by various hawkmoths (family Sphingidae). The flowers are remarkably similar in form and function to those of L. washingtonianum, which is also moth-pollinated, but this resemblance is due to evolutionary convergence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||
Key |
|
|||||||||||||||||
Source | FNA vol. 26, p. 188. | FNA vol. 26, p. 191. | ||||||||||||||||
Parent taxa | ||||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | L. parryi var. kessleri | |||||||||||||||||
Name authority | Kellogg: Hesperian (San Francisco) 3: 300. (1859) | S. Watson: Proc. Davenport Acad. Nat. Sci. 2: 188. (1878) | ||||||||||||||||
Web links |