Lilium pardalinum |
Lilium kelleyanum |
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California tiger lily, leopard lily, panther lily |
Kelley's lily |
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Bulbs | rhizomatous, usually branching, continuously scaly, 1.4–5.1 × 3.9–19 cm, 0.2–0.6 times taller than long; scales sometimes unsegmented but always some 2–4-segmented on each bulb, longest 1–3.3 cm; stem roots absent. |
rhizomatous, unbranched, continuously scaly, 1.4–3.1 × 3.8–8.2 cm, 0.3–0.6 times taller than long; scales 2–3-segmented, a few unsegmented, longest 0.8–2.7 cm; stem roots absent. |
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Stems | to 2.8 m, strongly clonal and thus forming dense colonies, to weakly clonal and forming small colonies or clumps. |
to 2.2 m. Buds rounded in cross section. |
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Buds | rounded in cross section. |
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Leaves | usually ± evenly distributed along stem, rarely concentrated proximally, scattered or in 1–6 whorls or partial whorls, 3–19 leaves per whorl, horizontal and drooping at tips to ascending, 4.9–26.5 × 0.3–5.6 cm, 3–34 times longer than wide; blade usually ± elliptic, wide or narrow, margins usually not undulate, apex acute, often narrowly so; veins and margins ± smooth abaxially. |
scattered or usually in 1–4 whorls or partial whorls, 3–10 leaves per whorl, horizontal and drooping at tips, 7.8–15.8 × 1.1–5.3 cm, 3.2–10.8 times longer than wide; blade usually elliptic, occasionally weakly lanceolate or oblanceolate, margins not undulate, apex narrowly acute; veins and margins ± smooth abaxially. |
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Inflorescences | racemose, 1–28(–35)-flowered. |
racemose, 1–15(–25)-flowered. |
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Flowers | pendent, usually not fragrant; perianth Turk’s-cap-shaped; sepals and petals reflexed 1/4–1/3 along length from base, yellow, yellow-orange, or orange proximally, darker orange to red-orange to red on distal 1/5–3/5 (entirely orange or yellow-orange in subsp. wigginsii), with maroon spots concentrated proximally and always surrounded by yellow or orange if extending into distal reddish zone, conspicuously green abaxially on proximal ± 1/5, not distinctly clawed; sepals not ridged abaxially, 3.5–10.4 × 0.9–2.2 cm; petals 3.4–10.2 × 0.9–2.5 cm; stamens moderately to strongly exserted; filaments moderately to widely spreading, diverging 7°–22° from axis; anthers ± magenta or sometimes orange, orange-pink, or pale yellow, 0.5–2.2 cm; pollen red-brown, red-orange, brown-orange, rust, orange, or yellow; pistil 3.1–7.5 cm; ovary 1–2.2 cm; style green, often pale, rarely sordid; pedicel 6–32 cm. |
pendent, mildly fragrant; perianth Turk’s-cap-shaped; sepals and petals reflexed 2/5 along length from base, yellow or yellow-orange, usually with copious maroon spots, not distinctly clawed; sepals not ridged abaxially, 4.3–5.7 × 0.9–1.5 cm; petals 4.2–5.7 × 0.9–1.5 cm; stamens moderately exserted; filaments moderately spreading, diverging 9°–19° from axis; anthers magenta or dull red, 0.3–0.6 cm; pollen pale orange, rust, orange, or tan-orange; pistil 2.6–3.4 cm; ovary 0.8–1.4 cm; style green; pedicel 8.5–20.7 cm. |
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Capsules | 2.2–5.7 × 1.2–2.1 cm, 1.5–3.7 times longer than wide. |
1.5–2.9 × 1–1.9 cm, 1.1–2 times longer than wide. |
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Seeds | 123–264. |
96–169. |
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2n | = 24. |
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Lilium pardalinum |
Lilium kelleyanum |
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Phenology | Flowering summer (Jul–Aug). | |||||||||||||||||
Habitat | Hillside seeps, wet thickets, along streams in coniferous forests | |||||||||||||||||
Elevation | 2200–2900 m (7200–9500 ft) | |||||||||||||||||
Distribution |
CA; OR
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CA
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Discussion | Subspecies 5 (5 in the flora). The subspecies of Lilium pardalinum display a classic pattern of discrete geographical ranges with intervening zones of introgression, and no two occur sympatrically without intermixing. Plants in the hybrid zones are confusing in appearance and cannot be assigned to subspecies. However, each subspecies is fairly well marked within its core distribution. With the exception of subsp. pitkinense, the subspecies of L. pardalinum can be common plants in the proper habitats within their rather narrow distributions. Leaf size and shape are quite variable in Lilium pardalinum subspecies and often clearly dependent on environment. In populations that typically have narrow, ascending leaves, shaded plants often have wide, horizontal leaves. This hampers taxonomic separation as well as identification, especially of herbarium specimens. Further field study is desirable. Lilium pardalinum is primarily pollinated by western tiger swallowtails (Papilio rutulus Lucas, family Papilionidae) and pale swallowtails (P. eurymedon Lucas); several species of hummingbirds (family Trochilidae) are also important visitors, especially when butterflies are rare. The Atsugewi, Karok, and Yana ate Lilium pardalinum bulbs steamed or baked in an earth oven (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lilium kelleyanum has in common with L. parvum a short, relatively wide fruit and small flowers, and divides with that species the montane and subalpine Sierra Nevada on both sides of the divide from Lake Tahoe south. The two species are parapatrically distributed with an intervening zone of introgression; intergrades occur, among other places, in the June Lake region about ten miles south of Mono Lake in Mono County and in adjacent Madera County. The two species appear to be sister taxa (M. W. Skinner 1988), which argues against subsuming L. kelleyanum as a subspecies of L. pardalinum, as has been suggested by several students of the group. Despite the close affinity of L. kelleyanum to L. parvum, taxonomic custom within the genus dictates that they be retained as separate species since floral features are well differentiated. Lilium kelleyanum is pollinated by swallowtail butterflies (J. S. Davis 1956). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 188. | FNA vol. 26, p. 187. | ||||||||||||||||
Parent taxa | ||||||||||||||||||
Sibling taxa | ||||||||||||||||||
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Synonyms | L. fresnense, L. inyoense, L. nevadense var. fresnense, L. nevadense var. monense, L. pardalinum var. parviflorum | |||||||||||||||||
Name authority | Kellogg: Hesperian (San Francisco) 3: 300. (1859) | Lemmon: Sierra Club Bull. 4: 300. (1903) | ||||||||||||||||
Web links |