Helenium autumnale |
Asteraceae tribe Heliantheae subtribe Gaillardiinae |
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common sneezeweed, fall sneezeweed, hélénie automnale, large-flower sneezeweed, mountain sneezeweed |
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Habit | Perennials, 50–130 cm. | Annuals, biennials, perennials, subshrubs, or shrubs, 3–160 cm. | ||||||||||||||||||||||||||||||||||||||||
Stems | 1(–7), branched distally, strongly winged, sparsely to densely hairy proximally, moderately to densely hairy distally. |
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Leaves | usually moderately to densely hairy, sometimes glabrous; basal blades (withered by flowering) lanceolate, oblanceolate, or obovate, entire or weakly lobed; proximal and mid blades obovate to oblanceolate, usually dentate or entire; distal blades oblanceolate to lanceolate, entire or dentate. |
basal, basal and cauline, or cauline; alternate; petiolate or sessile; blades mostly elliptic, filiform, lanceolate, linear, oblong, ovate, or spatulate, often 1–2-pinnately or -ternately lobed, ultimate margins entire or toothed, faces glabrous or hairy, usually gland-dotted. |
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Peduncles | 3–10 cm, moderately to densely hairy. |
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Involucres | globoid, 8–20 × 8–23 mm. |
mostly campanulate, cylindric, hemispheric, obconic, or ± rotate. |
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Receptacles | mostly conic, convex, flat, globose, hemispheric, or ovoid, epaleate (smooth or slightly to deeply pitted, sometimes with setiform enations, sometimes gland-dotted; outer disc florets rarely subtended by paleae in Amblyolepis). |
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Ray florets | 8–21, pistillate, fertile; corollas yellow, 10–23 × 4–10 mm. |
0, or 1–55 (in 1–3 series), pistillate and fertile, or styliferous and sterile, or neuter; corollas mostly yellow to orange, sometimes wholly or partly purple, red-brown, or reddish. |
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Disc florets | 200–400(–800+); corollas yellow proximally, yellow to yellow-brown distally, 2.4–4 mm, lobes 5. |
5–250+, usually bisexual and fertile, rarely functionally staminate; corollas yellow, orange, or wholly or partly purple, red-brown, or reddish, tubes much shorter than to about equaling slightly to abruptly and greatly dilated, funnelform to campanulate or urceolate throats, lobes 5, deltate to lance-deltate (equal or unequal, often attenuate, glabrous, papillate, or hairy, hairs sometimes moniliform); anther thecae pale; stigmatic papillae in 2 lines. |
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Phyllaries | (connate proximally) moderately to densely hairy. |
persistent, 5–50+ in 2–3+ series, distinct or outer connate, mostly elliptic, lanceolate, linear, oblanceolate, oblong, or ovate, subequal or unequal (erect or reflexed in fruit, usually herbaceous to scarious, margins seldom scarious). |
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Calyculi | 0. |
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Heads | 5–70(–100+) per plant, in paniculiform arrays. |
mostly radiate or discoid (rarely radiant), usually borne singly, sometimes in glomerules or corymbiform arrays. |
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Cypselae | 1–2 mm, sparsely to moderately hairy; pappi of 5–7 entire, aristate scales (0.5–)0.9–1.5(–1.8) mm. |
usually obpyramidal, sometimes clavate, columnar, or obconic (lengths usually 1–2.5, rarely to 3.5 diams.), mostly densely and shaggily hairy, sometimes sparsely hairy or glabrate; pappi usually persistent, of 2–12 scales, sometimes 0 or of 35–150 bristles (Psathyrotes). |
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2n | = 32, 34, 36. |
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Helenium autumnale |
Asteraceae tribe Heliantheae subtribe Gaillardiinae |
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Phenology | Flowering (Jul–)Aug–Oct(–Nov). | |||||||||||||||||||||||||||||||||||||||||
Habitat | Roadsides, fields, along streams, ditches, seepage areas, around ponds and lakes | |||||||||||||||||||||||||||||||||||||||||
Elevation | 20–2600 m (100–8500 ft) | |||||||||||||||||||||||||||||||||||||||||
Distribution |
AL; AR; AZ; CA; CO; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; WY; AB; BC; MB; NT; ON; QC; SK
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Mexico; South America; Mostly sw United States |
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Discussion | Genera 12, species 123 (11 genera, 73 species in the flora). The circumscription of Gaillardiinae adopted here is that of H. Robinson (1981). In studies of epaleate Heliantheae, B. G. Baldwin adopted a narrower circumscription of the subtribe (Baldwin and B. L. Wessa 2000; Baldwin et al. 2002). Gaillardiinae in sense of Baldwin includes Balduina, Gaillardia, and Helenium; dispositions by Baldwin of other genera of Gaillardiinae in the broad sense include: Plateilema in Plateileminae, Psathyrotes and Trichoptilium in Psathyrotinae, and Amblyolepis, Baileya, Hymenoxys, Psilostrophe, and Tetraneuris in Tetraneuriinae (all in Helenieae sensu Baldwin). Psilostrophinae B. L. Turner & A. M. Powell is a superfluous name (circumscription included type of Riddelliinae O. Hoffmann). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 431. | FNA vol. 21, p. 415. | ||||||||||||||||||||||||||||||||||||||||
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Synonyms | H. autumnale var. canaliculatum, H. autumnale var. fylesii, H. autumnale var. grandiflorum, H. autumnale var. montanum, H. autumnale var. parviflorum, H. latifolium, H. parviflorum | |||||||||||||||||||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 2: 886. (1753) | Lessing: Linnaea 6: 516. (1831) | ||||||||||||||||||||||||||||||||||||||||
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