Eurybia spinulosa |
Eurybia macrophylla |
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Apalachicola aster |
aster à grandes feuilles, bigleaf aster, large-leaf wood-aster |
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Habit | Plants 20–70 cm; solitary or clumped, eglandular; rhizomes short and stout or elongate and wiry, or caudices. | Plants 15–110 cm; in extensive, dense clones (with abundant, sterile rosettes); rhizomes branched, herbaceous. |
Stems | 1(–3+), erect, simple, ± villous to glabrescent. |
1, erect, simple, straight to ± flexuous, glabrous or sparsely villous, stipitate-glandular, more densely so distally (at least in arrays). |
Leaves | strongly basal and cauline, linear, firm, ± fleshy, margins indurate, ± revolute, entire to spinose-serrate, smooth to remotely scabridulous or ciliate, spines indurate, finely parallel-veined with evident midribs, apices acute, revolute-indurate, faces glabrescent (minute hairs bulbous at base, threadlike distally); basal and proximal cauline persistent, sessile or petiolate (narrowing between bases and blades), blades lance-linear to linear, 100–300 × (1–)2–5 mm, bases ± marcescent, sheathing, ciliate; cauline sessile, blades linear to lance-linear, 20–95 × 3–5 mm, progressively reduced distally, bases rounded- to auriculate-clasping, adaxial faces sparsely villous in distal, the distal subtending heads boat-shaped. |
basal and cauline, thick, firm, margins coarsely serrate (proximal) to crenate-serrate or entire (distal), scabrous, teeth mucronate, apices acuminate, sharply mucronate, abaxial faces glabrescent to sparsely strigose, veins stipitate-glandular, adaxial sparsely strigose, ± densely villous on veins, short-stipitate-glandular (more so on distal); basal withering by flowering, petioles (25–)80–170 mm, ± winged, bases sheathing, ciliate, blades broadly ovate, (45–)110–250 × (25–)55–150 mm, bases deeply cordate (sinuses curved), sometimes slightly oblique; cauline petiolate (3–110 mm), gradually more broadly winged and sometimes ± auriculate distally, ± clasping, ciliate, to subpetiolate or sessile, blades broadly ovate to ovate, 22–104 × 9–80 mm, gradually reduced distally, bases cordate to rounded or attenuate to cuneate; distal (arrays) sessile, blades ovate or broadly lanceolate or elliptic to obovate or oblanceolate, 9–38 × 3–13 mm, abruptly reduced distally, bases ± clasping to rounded or cuneate (arrays), margins crenate-serrate or entire. |
Peduncles | 0 (usually) or ascending, 1–17+ mm, sparsely villosulous; bracts 0–2, ascending, lanceolate, bases not indurate, rounded (boat-shaped), margins ciliate, faces glabrous. |
firm, 0–4 cm, densely stipitate-glandular; bracts 0–2, ovate-lanceolate, ± stipitate-glandular. |
Involucres | campanulate, 6.5–9.7 mm, shorter than pappi. |
broadly campanulate to campanulate, (6–)7–11 mm, shorter than pappi. |
Ray florets | 8–17; laminae pale purple to purplish white, 10–16(–20) × 1–1.8 mm. |
9–20; corollas ± deep lavender or violet to occasionally white, (7–)11–15 × 1.4–2.4 mm. |
Disc florets | 18–30; corollas yellow, 5.5–7.6 mm, barely ampliate, tubes much shorter than tubular-funnelform throats (1–2 mm), lobes erect, lanceolate, 0.65–1 mm. |
20–40; corollas cream-color or light yellow, becoming purple, 6–7.5 mm, slightly ampliate, tubes (ca. 4 mm) longer than funnelform throats (ca 1.2 mm), lobes reflexed, lanceolate, 1–1.6 mm. |
Phyllaries | 20–40 in 4–5 series, green, often ± involute in distal 1/2–2/3 (outer) to 1/3 (inner), densely nerved (nerves not thickened), lanceolate, unequal, coriaceous, bases indurate, rounded (outer), margins entire, indurate (outer) or scarious and often purplish (inner), sparsely ciliate, apices acute to acuminate, indurate, apiculate, adaxial faces glabrous or sparsely villosulous. |
32–35 in 5–6 series, appressed, often purple-tinged, usually obovate to oblong, seldom ovate (outer) to oblanceolate or linear-lanceolate (inner) (innermost to 7 mm), strongly unequal, bases indurate, keeled or rounded adaxially, dark green zones wide, 1/3–1/2 distal portion (rarely to base in outer) to less pronounced along midveins or none (inner), margins hyaline or reddish, scarious, ± erose, densely villoso-ciliate apically, apices obtuse to rounded or acute (inner), adaxial faces ± villous, ± stipitate-glandular. |
Heads | 3–16+ in spiciform to narrow, racemiform arrays. |
(2–)8–90+ in flat-topped, corymbiform arrays. |
Cypselae | brown to gray-brown, fusiform, ± compressed, 2–2.5 mm, ribs 7–10, faces ± strigillose; pappi of burnt-orange (coarse, sometimes apically clavellate) bristles 6–7.5 mm, as long as or slightly longer than disc corollas. |
brown, fusiform or cylindro-obconic, compressed, 2.6–4.5 mm, ribs 7–12 (golden brown), faces glabrous or sparsely strigillose distally; pappi of tawny to orangish (sometimes clavellate) bristles 5–7.5 mm, slightly longer than disc corollas. |
2n | = 72. |
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Eurybia spinulosa |
Eurybia macrophylla |
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Phenology | Flowering May–Jul. | Flowering late summer–early fall (late Aug–Oct). |
Habitat | Moist to dry, acid sandy peats, savannas in long-leaf pinelands, fire-maintained | Moist to dry soils, hemlock-northern hardwoods, beech-maple or pine forests, Appalachian spruce-fir forests, aspen, pine or open spruce woodlands, thickets, clearings, shaded roadsides |
Elevation | 0–50 m (0–200 ft) | 0–1300 m (0–4300 ft) |
Distribution |
FL |
CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; MB; NB; NS; ON; PE; QC [Introduced in n Europe]
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Discussion | Of conservation concern. Eurybia spinulosa is known only from the Apalachicola River drainage of the Florida panhandle; it is of conservation concern in Florida and is a facultative wetland indicator. Much of its habitat has now been lost to development (R. Kral 1983, vol. 2). Kral published a map of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eurybia macrophylla, a species of the northeastern deciduous or mixed forest, extends north into the southern boreal forest of Ontario-Quebec, and south along the Blue Ridges into northern Georgia. It becomes rare at the western edge of its range (Manitoba, Illinois, Iowa, Missouri). The plant has been reported from Mississippi but no voucher was seen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 381. | FNA vol. 20, p. 375. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Aster spinulosus, Heleastrum spinulosum | Aster macrophyllus, Aster macrophyllus var. apricensis, Aster macrophyllus var. excelsior, Aster macrophyllus var. ianthinus, Aster macrophyllus var. pinguifolius, Aster macrophyllus var. sejunctus, Aster macrophyllus var. velutinus, Aster multiformis, Aster riciniatus, Biotia latifolia, Biotia macrophylla, E. jussiei |
Name authority | (Chapman) G. L. Nesom: Phytologia 77: 262. (1995) | (Linnaeus) Cassini: in F. Cuvier, Dict. Sci. Nat. ed. 2, 37: 487. (1825) |
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