Eurybia radula |
Eurybia |
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aster rude, low rough aster, rough wood-aster |
aster |
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Habit | Plants 10–100 cm; in lax clones, eglandular; rhizomes elongate or short, filiform or cordlike, herbaceous, remotely scaly. | Perennials, 10–120 cm (rhizomes long and slender to short and thick, sometimes cormoid, often becoming woody). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | 1, erect, straight, often reddish, simple, glabrous or glabrescent to sparsely villous proximally, villous distally. |
ascending to erect, usually simple, rarely branched proximally, glabrous or ± densely hairy, usually eglandular, sometimes stipitate-glandular. |
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Leaves | cauline, rugose-veined, margins ± revolute, ± remotely, ± deeply serrate, scabrous, teeth ± indurate (mucronate), apices mucronulate, abaxial faces scabrous, adaxial ± villous, particularly on veins (increasingly so distally); proximalmost withering by flowering, winged-petiolate, petioles equaling or longer than blades, bases ± sheathing, sometimes marcescent, blades spatulate to oblanceolate, 10–42 × 3–8 mm, sometimes reduced, smaller than mid, bases attenuate to cuneate, apices obtuse to acute; mid sessile, lanceolate, oblanceolate, or narrowly obovate to narrowly elliptic or elliptic, (23–)30–115 × 5–36 mm, gradually reduced distally, bases tapering or rounded to ± auriculate-clasping (often narrowed above base), apices acute to acuminate; distal (arrays) sessile, (6–)10–40 × 2–7 mm. |
basal and cauline; alternate; sessile or petiolate; blades cordate, ovate, obovate, elliptic, or oblong to spatulate, oblanceolate, or lanceolate, usually gradually reduced distally, margins entire or serrate, sometimes spinulose-serrate, faces glabrate to hairy, usually eglandular, sometimes stipitate-glandular. |
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Peduncles | ± densely villous; bracts 0–2, sometimes subtending heads. |
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Involucres | campanulate, 5.5–9 mm, shorter than pappi. |
cylindro-campanulate to broadly campanulate, (4–14(–16) ×) 4–25+ mm. |
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Receptacles | flat to slightly convex, pitted, epaleate. |
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Ray florets | 13–30; corollas pale blue-violet, 10.5–15 × 1.4–2 mm. |
5–60, pistillate, fertile; corollas white to purple (coiling at maturity). |
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Disc florets | 37–72; corollas yellow, 4.5–6 mm, slightly ampliate, 4.8–5.8 mm, tube lengths about twice funnelform throats (2.7–3.4 mm), lobes erect, lanceolate, 0.6–1 mm. |
8–260, bisexual, fertile; corollas yellow, becoming purple at maturity, barely to abruptly ampliate, tubes shorter to longer than funnelform to campanulate throats, lobes 5, usually erect to spreading, sometimes ± reflexed, deltate, triangular, or lanceolate; style-branch appendages lanceolate. |
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Phyllaries | 30–50 in 4–5 series, oblong or oblong-lanceolate, seldom ovate (outer), lanceolate to linear-lanceolate (inner), strongly unequal, membranous, bases indurate, rounded (outer), green zones full-width, occasionally purplish (inner), in distal 1/2 (outer; rarely 2/3 and foliaceous) to less than 1/6 and paler or none (inner), margins hyaline (seldom purplish), narrowly scarious, ± erose, densely ciliate, apices usually appressed, often squarrose, usually acute, sometimes obtuse, mucronulate, faces glabrous or sparsely strigillose. |
20–140 in 3–7 series, 1-nerved (usually rounded adaxially, sometimes low-keeled), broadly ovate or oblong to oblanceolate, lanceolate, or linear, unequal, bases indurate (rarely wholly foliaceous), margins narrowly scarious (seldom herbaceous), often ciliolate (green zones ± basally truncate), in distal 1/3–3/4 of phyllary (outer) to less than 1/6 and only along midnerves (inner), apices obtuse to acute, faces glabrous, ± strigillose, puberulent, scabrellous, strigoso-villous, or villous, sometimes ± stipitate-glandular. |
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Heads | 1–25+ in open corymbiform arrays or borne singly. |
radiate, usually in corymbiform arrays, rarely borne singly. |
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Cypselae | tawny to stramineous, fusiform to cylindro-obconic, slightly compressed, 2.7–3.5 mm, ribs 8–18 (stramineous or reddish brown), ± crowded, faces glabrous; pappi of pale cinnamon bristles 4.5–5.2 mm, shorter than disc corollas. |
cylindro-obconic to fusiform, ± compressed, 7–12(–18)-nerved, faces glabrous or sparsely to densely strigillose, eglandular; pappi persistent, of 35–70+, reddish, orange, cinnamon, tawny, tan, yellowish, or pinkish, unequal, soft to stiff, barbellate or barbellulate, often apically ± clavate bristles in 2–4 series. |
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x | = 9. |
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2n | = 18. |
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Eurybia radula |
Eurybia |
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Phenology | Flowering late summer–early fall. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Fens, sphagnum bogs, lake and creek shores, edges of or openings in wet spruce or tamarack forests, open boggy woods, ± wet meadows, ditches | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0–600+ m (0–2000+ ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CT; DC; DE; KY; MA; MD; ME; NH; NJ; NY; PA; RI; VA; VT; WV; NB; NL; NS; ON; PE; QC; SPM
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North America; n Eurasia |
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Discussion | Eurybia radula reaches its northwestern limit south of James Bay in northeastern Ontario, where it is rare (J. C. Semple et al. 2002). It is vulnerable in most of its United States range. Aster radula var. strictus (Pursh) A. Gray (for which no combination exists in Eurybia) appears to be a reduced, more slender plant with 1–8 heads (M. L. Fernald 1950) from more northern parts of the range (Newfoundland and Labrador, northern Quebec, northern Nova Scotia), presumably due to harsher conditions; an entire spectrum of intermediates appears to exist with the typical variety to the south, however, and it is unclear whether the variety should be recognized. It seems that the common name, rough aster, stems from a misinterpretation of the Latin epithet radula, which means scraper (rough would be radulans). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 23 (23, including 1 hybrid, in the flora). Eurybia traditionally has been treated within Aster in a broad sense. G. L. Nesom (1994b), in his review of North American asters, showed that Aster in a broad sense does not form a natural group and proposed splitting it into several genera, among which Eurybia is one. In his treatment, Nesom included Herrickia within Eurybia, as sect. Herrickia in subg. Eurybia. Such views were generally supported in molecular phylogenetic studies (J. C. Semple et al. 2002). L. Brouillet et al. (2004) showed, however, that Oreostemma, Herrickia, Eurybia, and Triniteurybia form a grade at the base of the Machaerantherinae and that Herrickia and Eurybia are distinct. The subgenera and sections proposed by G. L. Nesom (1994b), based on anterior taxonomy, could not be confirmed in the molecular studies cited above. I chose not to use subgeneric limits as proposed by Nesom because they may not reflect actual relationships. For instance, there is a clear gradation between members of sect. Calliastrum (Torrey & A. Gray) G. L. Nesom (subg. Eurybia) and sect. Heleastrum (de Candolle) G. L. Nesom. Also, I do not recognize sect. Eryngiifoliae (Alexander) G. L. Nesom distinct from sect. Heleastrum, as there is no clear demarcation between the two as currently defined. Finally, sect. Radulini (Rydberg) G. L. Nesom appears artificial to me, but currently there is no good way to reassign its species. The Eurybia radulina complex of western North America clearly constitute a group, but it is unclear whether the western E. conspicua or the eastern E. radula and E. saxicastelli are close to them. Members of other sections may have played a role in the reticulate evolution of sect. Eurybia, even though it is well marked by its cordate leaves and disc florets with long tubes and short, campanulate corollas. Therefore, species are described below in a rough taxonomic order, with diploids listed before polyploids of the same group. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 371. | FNA vol. 20, p. 365. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Aster radula, Aster radula var. strictus | Aster subg. E. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Aiton) G. L. Nesom: Phytologia 77: 261. (1995) | (Cassini) Cassini: in F. Cuvier, Dict. Sci. Nat. ed. 2, 16: 46. (1820) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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