Eurybia |
Eurybia integrifolia |
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aster |
thick-stem aster |
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Habit | Perennials, 10–120 cm (rhizomes long and slender to short and thick, sometimes cormoid, often becoming woody). | Plants 15–70 cm, usually in clumps, sometimes in large clones, densely long-stipitate-glandular distally; woody, branched, thick, usually short rhizomes or short caudices. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | ascending to erect, usually simple, rarely branched proximally, glabrous or ± densely hairy, usually eglandular, sometimes stipitate-glandular. |
1–3+, straight, stout, glabrous or sparsely hispid proximally, distally ± hispido-villous. |
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Leaves | basal and cauline; alternate; sessile or petiolate; blades cordate, ovate, obovate, elliptic, or oblong to spatulate, oblanceolate, or lanceolate, usually gradually reduced distally, margins entire or serrate, sometimes spinulose-serrate, faces glabrate to hairy, usually eglandular, sometimes stipitate-glandular. |
basal and cauline, firm, margins entire, strigoso-ciliate, distal also stipitate-glandular, apices mucronate, faces glabrous or glabrescent to ± densely hispid or strigose (then ± scabrous), particularly on veins, midveins sometimes notably hispido-villous, proximally ± sparsely, distally ± densely stipitate-glandular; basal and proximal cauline long-petiolate (to 100+ mm), petioles ± broadly winged, bases sheathing or auriculate-clasping, blades ovate-lanceolate to narrowly elliptic or oblanceolate, 33–180+ × 11–50 mm, bases attenuate, apices acute or obtuse to rounded; mid sessile, blades oblong or oblanceolate to oblong-lanceolate, lanceolate, or lance-ovate, 30–140 × 7–27 mm, gradually reduced distally, bases auriculate-clasping, apices usually acute, rarely obtuse; distal (arrays) narrowly ovate to lanceolate, 9–50 × 3–20 mm. |
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Peduncles | ± densely long-stipitate-glandular; bracts 0(–2), densely stipitate-glandular. |
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Involucres | cylindro-campanulate to broadly campanulate, (4–14(–16) ×) 4–25+ mm. |
campanulate, 8–14 mm, much shorter than pappi. |
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Receptacles | flat to slightly convex, pitted, epaleate. |
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Ray florets | 5–60, pistillate, fertile; corollas white to purple (coiling at maturity). |
8–27; corollas violet-purple, 10–15 × 1.2–2.2 mm. |
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Disc florets | 8–260, bisexual, fertile; corollas yellow, becoming purple at maturity, barely to abruptly ampliate, tubes shorter to longer than funnelform to campanulate throats, lobes 5, usually erect to spreading, sometimes ± reflexed, deltate, triangular, or lanceolate; style-branch appendages lanceolate. |
20–50; corollas pale yellow turning pinkish or purplish, 6–7.8 mm, slightly ampliate, tubes much shorter than cylindric to narrowly funnelform throats, lobes erect, lanceolate, 0.6–0.8 mm. |
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Phyllaries | 20–140 in 3–7 series, 1-nerved (usually rounded adaxially, sometimes low-keeled), broadly ovate or oblong to oblanceolate, lanceolate, or linear, unequal, bases indurate (rarely wholly foliaceous), margins narrowly scarious (seldom herbaceous), often ciliolate (green zones ± basally truncate), in distal 1/3–3/4 of phyllary (outer) to less than 1/6 and only along midnerves (inner), apices obtuse to acute, faces glabrous, ± strigillose, puberulent, scabrellous, strigoso-villous, or villous, sometimes ± stipitate-glandular. |
25–40 in 3–4 series, inner often purplish, oblong-lanceolate (outer) to linear-lanceolate (inner), ± unequal, membranous, bases pale, indurate, sometimes rounded (outer), distally foliaceous (3/4+ in outer, seldom to base, to 1/5 in inner) and wider than bases, margins narrowly scarious (non-foliaceous parts), purplish (at least inner), ciliate and/or stipitate-glandular (along foliaceous parts), apices squarrose, usually acute, sometimes acuminate, faces densely stipitate-glandular. |
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Heads | radiate, usually in corymbiform arrays, rarely borne singly. |
3–41+ in elongate, racemo-corymbiform arrays, branches ascending. |
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Cypselae | cylindro-obconic to fusiform, ± compressed, 7–12(–18)-nerved, faces glabrous or sparsely to densely strigillose, eglandular; pappi persistent, of 35–70+, reddish, orange, cinnamon, tawny, tan, yellowish, or pinkish, unequal, soft to stiff, barbellate or barbellulate, often apically ± clavate bristles in 2–4 series. |
greenish stramineous, fusiform-obconic, slightly compressed, 4.2–4.7 mm, ribs 7–10, faces ± densely hirtellous; pappi of stramineous to tawny bristles 7–8 mm, ± equaling disc corollas. |
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x | = 9. |
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2n | = 18. |
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Eurybia |
Eurybia integrifolia |
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Phenology | Flowering summer–early fall. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Drier meadows, open, moist woodlands, in sedge-willow, sagebrush, Douglas fir, and spruce communities | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1600–3200 m (5200–10500 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
North America; n Eurasia |
CA; ID; MT; NV; OR; UT; WA; WY
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Discussion | Species 23 (23, including 1 hybrid, in the flora). Eurybia traditionally has been treated within Aster in a broad sense. G. L. Nesom (1994b), in his review of North American asters, showed that Aster in a broad sense does not form a natural group and proposed splitting it into several genera, among which Eurybia is one. In his treatment, Nesom included Herrickia within Eurybia, as sect. Herrickia in subg. Eurybia. Such views were generally supported in molecular phylogenetic studies (J. C. Semple et al. 2002). L. Brouillet et al. (2004) showed, however, that Oreostemma, Herrickia, Eurybia, and Triniteurybia form a grade at the base of the Machaerantherinae and that Herrickia and Eurybia are distinct. The subgenera and sections proposed by G. L. Nesom (1994b), based on anterior taxonomy, could not be confirmed in the molecular studies cited above. I chose not to use subgeneric limits as proposed by Nesom because they may not reflect actual relationships. For instance, there is a clear gradation between members of sect. Calliastrum (Torrey & A. Gray) G. L. Nesom (subg. Eurybia) and sect. Heleastrum (de Candolle) G. L. Nesom. Also, I do not recognize sect. Eryngiifoliae (Alexander) G. L. Nesom distinct from sect. Heleastrum, as there is no clear demarcation between the two as currently defined. Finally, sect. Radulini (Rydberg) G. L. Nesom appears artificial to me, but currently there is no good way to reassign its species. The Eurybia radulina complex of western North America clearly constitute a group, but it is unclear whether the western E. conspicua or the eastern E. radula and E. saxicastelli are close to them. Members of other sections may have played a role in the reticulate evolution of sect. Eurybia, even though it is well marked by its cordate leaves and disc florets with long tubes and short, campanulate corollas. Therefore, species are described below in a rough taxonomic order, with diploids listed before polyploids of the same group. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eurybia integrifolia is found in mountain ranges bordering the Basin and Range Province, from the Sierra Nevada and Cascade ranges in the west to the Rocky Mountains and Colorado Plateau in the east. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 365. | FNA vol. 20, p. 368. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Aster subg. E. | Aster integrifolius, Aster amplexifolius | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Cassini) Cassini: in F. Cuvier, Dict. Sci. Nat. ed. 2, 16: 46. (1820) | (Nuttall) G. L. Nesom: Phytologia 77: 260. (1995) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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