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Philadelphia daisy, Philadelphia fleabane, vergerette de philadelphie

branch fleabane, branching daisy, club-fruit willowherb, common eastern fleabane, daisy fleabane, plains fleabane, prairie fleabane, purple-leaf willowherb, rough fleabane, rough-stem daisy, rough-stem fleabane, vergerette rude, whitetop fleabane

Habit Annuals, biennials, or short-lived perennials, 4–80 cm; fibrous-rooted, caudices simple. Annuals, biennials, or short-lived perennials, 30–70 cm; fibrous-rooted, caudices simple, sometimes lignescent, sometimes producing rhizomes that bear leaf tufts at upturned ends.
Stems

erect (green proximally, leafy to arrays), hirsuto-villous to villous proximally, loosely strigose to sparsely hirsute distally, minutely glandular.

erect or ascending, sparsely to moderately strigose to strigillose or hirsuto-strigillose (hairs usually ascending, rarely spreading, 0.1–1.2 mm), eglandular.

Leaves

basal (persistent or withering by flowering) and cauline;

basal blades oblanceolate to obovate, (15–)30–110(–150) × 10–25(–40) mm, margins shallowly crenate to coarsely serrate or pinnately lobed, faces sparsely hirsute to villous, eglandular;

cauline blades oblong-oblanceolate to lanceolate, gradually reduced distally (bases clasping to auriculate-clasping).

basal (usually persistent through flowering) and cauline;

basal blades spatulate to broadly or narrowly oblanceolate to linear, (10–)30–150(–170) × 5–15(–21) mm;

cauline usually gradually reduced distally, continuing to near heads, margins entire or shallowly to deeply serrate or crenate, faces glabrous or glabrate to sparsely strigose or strigoso-hirsute, eglandular.

Involucres

4–6 × 6–15 mm.

(2–)3–4 × 5–12 mm.

Ray florets

150–250(–400);

corollas usually white, sometimes pinkish, 5–10 mm, laminae not coiling or tardily coiling.

50–100;

corollas white, less commonly pinkish or bluish, 4–6 mm, laminae coiling.

Disc corollas

2.1–3.2 mm.

1.5–2.5 mm (throats sometimes slightly indurate and inflated).

Phyllaries

in 2–3 series (sometimes basally connate), hirsuto-villous to sparsely hirsute or glabrous, sometimes minutely glandular.

in 2–4 series, glabrous, strigose, or sparsely hirsute, sometimes minutely glandular.

Heads

(1–)3–35 usually in corymbiform arrays (ultimate branches arising near stem tips).

10–200+ in loosely corymbiform to paniculiform-corymbiform arrays (on distal branches).

Cypselae

0.6–1.1 mm, 2-nerved, faces sparsely strigose;

pappi: outer of setae, inner of 15–20(–30) bristles.

(0.5–)0.9–1.2 mm, 2-nerved, faces sparsely strigose;

pappi: outer crowns of setae or scales, inner 0 (rays) or of 8–15 bristles (disc).

Erigeron philadelphicus

Erigeron strigosus

Distribution
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT [Introduced worldwide in temperate areas as a weed]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK [Introduced in Europe, Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf faces sparsely hirsute to villous
var. philadelphicus
1. Cauline leaf faces glabrous or glabrate
→ 2
2. Leaves of basal rosettes withering by flowering; salt marshes, British Columbia
var. glaber
2. Leaves of basal rosettes persistent through flowering; crevices in shale or dolomite, usually beside rivers or lakes, Ontario, Quebec, New York, and Vermont
var. provancheri
1. Phyllary hairs flattened, 0.5–1.2 mm; stem hairs appressed to spreading, 0.5–1 mm
var. septentrionalis
1. Phyllary hairs terete, mostly 0.1–0.5 mm; stem hairs appressed to ascending, 0.1–0.4(–0.8) mm
→ 2
2. Annual or rarely biennial, without rhizomes; various habitats, often ruderal
var. strigosus
2. Plants usually perennial (sometimes killed prematurely by severe drought), with rhizomes that bear overwintering leaf tufts at upturned ends; shallow soils over calcareous rock
→ 3
3. Basal leaves linear to linear-oblanceolate, 1–3.5(–6) mm wide; cauline leaf faces sparsely to moderately strigillose
var. dolomiticola
3. Basal leaves oblanceolate to narrowly obovate or spatulate, (3.2–)3.8–15(–21) mm wide; cauline leaf faces glabrous, margins and midveins ciliate
var. calcicola
Source FNA vol. 20, p. 319. FNA vol. 20, p. 345.
Parent taxa Asteraceae > tribe Astereae > Erigeron Asteraceae > tribe Astereae > Erigeron
Sibling taxa
E. abajoensis, E. acomanus, E. acris, E. aequifolius, E. algidus, E. aliceae, E. allocotus, E. alpiniformis, E. anchana, E. annuus, E. aphanactis, E. arenarioides, E. argentatus, E. arisolius, E. arizonicus, E. asperugineus, E. aureus, E. barbellulatus, E. basalticus, E. bellidiastrum, E. bigelovii, E. biolettii, E. blochmaniae, E. bloomeri, E. breweri, E. caespitosus, E. calvus, E. canaani, E. canus, E. cascadensis, E. cavernensis, E. cervinus, E. chrysopsidis, E. clokeyi, E. compactus, E. compositus, E. concinnus, E. consimilis, E. corymbosus, E. coulteri, E. cronquistii, E. davisii, E. decumbens, E. denalii, E. disparipilus, E. divergens, E. eatonii, E. elatior, E. elatus, E. elegantulus, E. elmeri, E. engelmannii, E. evermannii, E. eximius, E. filifolius, E. flabellifolius, E. flagellaris, E. flettii, E. foliosus, E. formosissimus, E. garrettii, E. geiseri, E. glabellus, E. glacialis, E. glaucus, E. goodrichii, E. gracilis, E. grandiflorus, E. greenei, E. heliographis, E. hessii, E. howellii, E. humilis, E. hyperboreus, E. hyssopifolius, E. inornatus, E. jonesii, E. kachinensis, E. karvinskianus, E. klamathensis, E. kuschei, E. lackschewitzii, E. lanatus, E. lassenianus, E. latus, E. leibergii, E. leiomerus, E. lemmonii, E. linearis, E. lobatus, E. lonchophyllus, E. maguirei, E. mancus, E. maniopotamicus, E. mariposanus, E. melanocephalus, E. miser, E. modestus, E. muirii, E. multiceps, E. nanus, E. nauseosus, E. nematophyllus, E. neomexicanus, E. nivalis, E. ochroleucus, E. oreganus, E. oreophilus, E. ovinus, E. oxyphyllus, E. pallens, E. parishii, E. parryi, E. peregrinus, E. petrophilus, E. pinnatisectus, E. piperianus, E. piscaticus, E. poliospermus, E. porsildii, E. pringlei, E. procumbens, E. pulchellus, E. pulcherrimus, E. pumilus, E. purpuratus, E. pygmaeus, E. quercifolius, E. radicatus, E. reductus, E. religiosus, E. rhizomatus, E. robustior, E. rybius, E. rydbergii, E. salishii, E. salmonensis, E. sanctarum, E. saxatilis, E. sceptrifer, E. scopulinus, E. serpentinus, E. sionis, E. sivinskii, E. sparsifolius, E. speciosus, E. strigosus, E. subglaber, E. subtrinervis, E. supplex, E. tenellus, E. tener, E. tenuis, E. tracyi, E. trifidus, E. tweedyi, E. uintahensis, E. uncialis, E. uniflorus, E. untermannii, E. ursinus, E. utahensis, E. vagus, E. velutipes, E. vernus, E. versicolor, E. vetensis, E. vicinus, E. vreelandii, E. watsonii, E. wilkenii, E. yukonensis
E. abajoensis, E. acomanus, E. acris, E. aequifolius, E. algidus, E. aliceae, E. allocotus, E. alpiniformis, E. anchana, E. annuus, E. aphanactis, E. arenarioides, E. argentatus, E. arisolius, E. arizonicus, E. asperugineus, E. aureus, E. barbellulatus, E. basalticus, E. bellidiastrum, E. bigelovii, E. biolettii, E. blochmaniae, E. bloomeri, E. breweri, E. caespitosus, E. calvus, E. canaani, E. canus, E. cascadensis, E. cavernensis, E. cervinus, E. chrysopsidis, E. clokeyi, E. compactus, E. compositus, E. concinnus, E. consimilis, E. corymbosus, E. coulteri, E. cronquistii, E. davisii, E. decumbens, E. denalii, E. disparipilus, E. divergens, E. eatonii, E. elatior, E. elatus, E. elegantulus, E. elmeri, E. engelmannii, E. evermannii, E. eximius, E. filifolius, E. flabellifolius, E. flagellaris, E. flettii, E. foliosus, E. formosissimus, E. garrettii, E. geiseri, E. glabellus, E. glacialis, E. glaucus, E. goodrichii, E. gracilis, E. grandiflorus, E. greenei, E. heliographis, E. hessii, E. howellii, E. humilis, E. hyperboreus, E. hyssopifolius, E. inornatus, E. jonesii, E. kachinensis, E. karvinskianus, E. klamathensis, E. kuschei, E. lackschewitzii, E. lanatus, E. lassenianus, E. latus, E. leibergii, E. leiomerus, E. lemmonii, E. linearis, E. lobatus, E. lonchophyllus, E. maguirei, E. mancus, E. maniopotamicus, E. mariposanus, E. melanocephalus, E. miser, E. modestus, E. muirii, E. multiceps, E. nanus, E. nauseosus, E. nematophyllus, E. neomexicanus, E. nivalis, E. ochroleucus, E. oreganus, E. oreophilus, E. ovinus, E. oxyphyllus, E. pallens, E. parishii, E. parryi, E. peregrinus, E. petrophilus, E. philadelphicus, E. pinnatisectus, E. piperianus, E. piscaticus, E. poliospermus, E. porsildii, E. pringlei, E. procumbens, E. pulchellus, E. pulcherrimus, E. pumilus, E. purpuratus, E. pygmaeus, E. quercifolius, E. radicatus, E. reductus, E. religiosus, E. rhizomatus, E. robustior, E. rybius, E. rydbergii, E. salishii, E. salmonensis, E. sanctarum, E. saxatilis, E. sceptrifer, E. scopulinus, E. serpentinus, E. sionis, E. sivinskii, E. sparsifolius, E. speciosus, E. subglaber, E. subtrinervis, E. supplex, E. tenellus, E. tener, E. tenuis, E. tracyi, E. trifidus, E. tweedyi, E. uintahensis, E. uncialis, E. uniflorus, E. untermannii, E. ursinus, E. utahensis, E. vagus, E. velutipes, E. vernus, E. versicolor, E. vetensis, E. vicinus, E. vreelandii, E. watsonii, E. wilkenii, E. yukonensis
Subordinate taxa
E. philadelphicus var. glaber, E. philadelphicus var. philadelphicus, E. philadelphicus var. provancheri
E. strigosus var. calcicola, E. strigosus var. dolomiticola, E. strigosus var. septentrionalis, E. strigosus var. strigosus
Synonyms E. annuus subsp. strigosus
Name authority Linnaeus: Sp. Pl. 2: 863. (1753) Muhlenberg ex Willdenow: Sp. Pl. 3: 1956. (1803)
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