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Philadelphia daisy, Philadelphia fleabane, vergerette de philadelphie

plains fleabane

Habit Annuals, biennials, or short-lived perennials, 4–80 cm; fibrous-rooted, caudices simple. Perennials (sometimes flowering early and appearing annual or biennial), 8–40 cm; usually taprooted (taproots woody or lignescent, early season forms sometimes fibrous-rooted), caudices usually woody, sometimes branched.
Stems

erect (green proximally, leafy to arrays), hirsuto-villous to villous proximally, loosely strigose to sparsely hirsute distally, minutely glandular.

(often multiple from bases; of previous year often persistent) ascending to spreading (proximally reddish in early season, sometimes proximally woody or lignescent), loosely strigose to villous (proximal hairs sometimes spreading), often sparsely glandular.

Leaves

basal (persistent or withering by flowering) and cauline;

basal blades oblanceolate to obovate, (15–)30–110(–150) × 10–25(–40) mm, margins shallowly crenate to coarsely serrate or pinnately lobed, faces sparsely hirsute to villous, eglandular;

cauline blades oblong-oblanceolate to lanceolate, gradually reduced distally (bases clasping to auriculate-clasping).

basal (often withering by late flowering) and cauline;

blades oblanceolate to spatulate, 20–50(–100) × 3–7(–18) mm, gradually reduced distally, margins entire or with 1–2(–3) pairs of teeth, faces loosely strigose (hairs ascending), eglandular.

Involucres

4–6 × 6–15 mm.

3–4.5 × 6–9(–12) mm.

Ray florets

150–250(–400);

corollas usually white, sometimes pinkish, 5–10 mm, laminae not coiling or tardily coiling.

24–65(–170);

corollas white, often with abaxial midstripe, often drying lilac, 4–7(–11) mm, laminae not coiling or reflexing.

Disc corollas

2.1–3.2 mm.

1.7–3.1 mm (throats indurate and slightly inflated).

Phyllaries

in 2–3 series (sometimes basally connate), hirsuto-villous to sparsely hirsute or glabrous, sometimes minutely glandular.

in 2–3 series, sparsely to moderately hirsute, minutely glandular.

Heads

(1–)3–35 usually in corymbiform arrays (ultimate branches arising near stem tips).

1(–3+, –50 in later season, branches at first from midstem or proximally, later more distal).

Cypselae

0.6–1.1 mm, 2-nerved, faces sparsely strigose;

pappi: outer of setae, inner of 15–20(–30) bristles.

1.1–1.4 mm, 2-nerved, faces sparsely strigose;

pappi: outer of setae, inner of (5–)12–17 bristles.

2n

= 27, 36.

Erigeron philadelphicus

Erigeron modestus

Phenology Flowering (Mar–)Apr–Jun(–Oct).
Habitat Rocky or gravelly sites, sand, clay, limestone, granite, sometimes deep sand, often with oak or oak-juniper, shrubland
Elevation 200–1700(–2300) m (700–5600(–7500) ft)
Distribution
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT [Introduced worldwide in temperate areas as a weed]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; KS; NM; OK; TX; Mexico (Coahuila, Chihuahua, Nuevo Leon)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Erigeron modestus is a polyploid complex apparently incorporating genes from E. flagellaris, E. tracyi, elements of E divergens, and the Mexican E. pubescens Kunth; arbitrary identifications may be necessary. In most of its range in Texas, E. modestus is not sympatric with any of those other taxa. Forms flowering in early season (Mar–Apr) often have stems red at the bases, dentate basal leaves 30–60(–100) × 5–10(–18) mm, and single, relatively large heads (involucres 9–12 mm diam, ray florets 50–170, corollas 6.5–11 mm) on ebracteate peduncles 5–15 cm. Later season plants have nearly intricately branched stems with more numerous, much smaller, diffusely arranged heads.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf faces sparsely hirsute to villous
var. philadelphicus
1. Cauline leaf faces glabrous or glabrate
→ 2
2. Leaves of basal rosettes withering by flowering; salt marshes, British Columbia
var. glaber
2. Leaves of basal rosettes persistent through flowering; crevices in shale or dolomite, usually beside rivers or lakes, Ontario, Quebec, New York, and Vermont
var. provancheri
Source FNA vol. 20, p. 319. FNA vol. 20, p. 341.
Parent taxa Asteraceae > tribe Astereae > Erigeron Asteraceae > tribe Astereae > Erigeron
Sibling taxa
E. abajoensis, E. acomanus, E. acris, E. aequifolius, E. algidus, E. aliceae, E. allocotus, E. alpiniformis, E. anchana, E. annuus, E. aphanactis, E. arenarioides, E. argentatus, E. arisolius, E. arizonicus, E. asperugineus, E. aureus, E. barbellulatus, E. basalticus, E. bellidiastrum, E. bigelovii, E. biolettii, E. blochmaniae, E. bloomeri, E. breweri, E. caespitosus, E. calvus, E. canaani, E. canus, E. cascadensis, E. cavernensis, E. cervinus, E. chrysopsidis, E. clokeyi, E. compactus, E. compositus, E. concinnus, E. consimilis, E. corymbosus, E. coulteri, E. cronquistii, E. davisii, E. decumbens, E. denalii, E. disparipilus, E. divergens, E. eatonii, E. elatior, E. elatus, E. elegantulus, E. elmeri, E. engelmannii, E. evermannii, E. eximius, E. filifolius, E. flabellifolius, E. flagellaris, E. flettii, E. foliosus, E. formosissimus, E. garrettii, E. geiseri, E. glabellus, E. glacialis, E. glaucus, E. goodrichii, E. gracilis, E. grandiflorus, E. greenei, E. heliographis, E. hessii, E. howellii, E. humilis, E. hyperboreus, E. hyssopifolius, E. inornatus, E. jonesii, E. kachinensis, E. karvinskianus, E. klamathensis, E. kuschei, E. lackschewitzii, E. lanatus, E. lassenianus, E. latus, E. leibergii, E. leiomerus, E. lemmonii, E. linearis, E. lobatus, E. lonchophyllus, E. maguirei, E. mancus, E. maniopotamicus, E. mariposanus, E. melanocephalus, E. miser, E. modestus, E. muirii, E. multiceps, E. nanus, E. nauseosus, E. nematophyllus, E. neomexicanus, E. nivalis, E. ochroleucus, E. oreganus, E. oreophilus, E. ovinus, E. oxyphyllus, E. pallens, E. parishii, E. parryi, E. peregrinus, E. petrophilus, E. pinnatisectus, E. piperianus, E. piscaticus, E. poliospermus, E. porsildii, E. pringlei, E. procumbens, E. pulchellus, E. pulcherrimus, E. pumilus, E. purpuratus, E. pygmaeus, E. quercifolius, E. radicatus, E. reductus, E. religiosus, E. rhizomatus, E. robustior, E. rybius, E. rydbergii, E. salishii, E. salmonensis, E. sanctarum, E. saxatilis, E. sceptrifer, E. scopulinus, E. serpentinus, E. sionis, E. sivinskii, E. sparsifolius, E. speciosus, E. strigosus, E. subglaber, E. subtrinervis, E. supplex, E. tenellus, E. tener, E. tenuis, E. tracyi, E. trifidus, E. tweedyi, E. uintahensis, E. uncialis, E. uniflorus, E. untermannii, E. ursinus, E. utahensis, E. vagus, E. velutipes, E. vernus, E. versicolor, E. vetensis, E. vicinus, E. vreelandii, E. watsonii, E. wilkenii, E. yukonensis
E. abajoensis, E. acomanus, E. acris, E. aequifolius, E. algidus, E. aliceae, E. allocotus, E. alpiniformis, E. anchana, E. annuus, E. aphanactis, E. arenarioides, E. argentatus, E. arisolius, E. arizonicus, E. asperugineus, E. aureus, E. barbellulatus, E. basalticus, E. bellidiastrum, E. bigelovii, E. biolettii, E. blochmaniae, E. bloomeri, E. breweri, E. caespitosus, E. calvus, E. canaani, E. canus, E. cascadensis, E. cavernensis, E. cervinus, E. chrysopsidis, E. clokeyi, E. compactus, E. compositus, E. concinnus, E. consimilis, E. corymbosus, E. coulteri, E. cronquistii, E. davisii, E. decumbens, E. denalii, E. disparipilus, E. divergens, E. eatonii, E. elatior, E. elatus, E. elegantulus, E. elmeri, E. engelmannii, E. evermannii, E. eximius, E. filifolius, E. flabellifolius, E. flagellaris, E. flettii, E. foliosus, E. formosissimus, E. garrettii, E. geiseri, E. glabellus, E. glacialis, E. glaucus, E. goodrichii, E. gracilis, E. grandiflorus, E. greenei, E. heliographis, E. hessii, E. howellii, E. humilis, E. hyperboreus, E. hyssopifolius, E. inornatus, E. jonesii, E. kachinensis, E. karvinskianus, E. klamathensis, E. kuschei, E. lackschewitzii, E. lanatus, E. lassenianus, E. latus, E. leibergii, E. leiomerus, E. lemmonii, E. linearis, E. lobatus, E. lonchophyllus, E. maguirei, E. mancus, E. maniopotamicus, E. mariposanus, E. melanocephalus, E. miser, E. muirii, E. multiceps, E. nanus, E. nauseosus, E. nematophyllus, E. neomexicanus, E. nivalis, E. ochroleucus, E. oreganus, E. oreophilus, E. ovinus, E. oxyphyllus, E. pallens, E. parishii, E. parryi, E. peregrinus, E. petrophilus, E. philadelphicus, E. pinnatisectus, E. piperianus, E. piscaticus, E. poliospermus, E. porsildii, E. pringlei, E. procumbens, E. pulchellus, E. pulcherrimus, E. pumilus, E. purpuratus, E. pygmaeus, E. quercifolius, E. radicatus, E. reductus, E. religiosus, E. rhizomatus, E. robustior, E. rybius, E. rydbergii, E. salishii, E. salmonensis, E. sanctarum, E. saxatilis, E. sceptrifer, E. scopulinus, E. serpentinus, E. sionis, E. sivinskii, E. sparsifolius, E. speciosus, E. strigosus, E. subglaber, E. subtrinervis, E. supplex, E. tenellus, E. tener, E. tenuis, E. tracyi, E. trifidus, E. tweedyi, E. uintahensis, E. uncialis, E. uniflorus, E. untermannii, E. ursinus, E. utahensis, E. vagus, E. velutipes, E. vernus, E. versicolor, E. vetensis, E. vicinus, E. vreelandii, E. watsonii, E. wilkenii, E. yukonensis
Subordinate taxa
E. philadelphicus var. glaber, E. philadelphicus var. philadelphicus, E. philadelphicus var. provancheri
Synonyms E. lobatus var. warnockii, E. plateauensis, E. warnockii
Name authority Linnaeus: Sp. Pl. 2: 863. (1753) A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 68. (1849)
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