Erigeron grandiflorus |
Erigeron divergens |
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large-flower daisy, large-flower fleabane, onestem fleabane, Rocky Mountain alpine fleabane |
branching fleabane diffuse daisy, diffuse daisy, diffuse fleabane, spreading fleabane |
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Habit | Perennials, 2–25 cm; rhizomatous, fibrous-rooted, caudices or rhizomes crownlike or branches relatively short and thick. | Annuals or short-lived perennials, (7–)12–40(–70) cm; taprooted, caudices simple if perennial. |
Stems | erect to decumbent-ascending, sparsely to moderately pilose to villoso-hirsute, often stipitate-glandular over all or part. |
(single or multiple from bases) erect to ascending, densely and evenly puberulous-hirsutulous (hairs spreading to spreading-descending or -ascending, often crinkly, bases not thickened), minutely glandular at least distally, usually throughout. |
Leaves | basal (persistent) and cauline (petioles equaling or shorter than blades); blades oblanceolate to obovate or spatulate, 10–60(–90) × 3–8(–14) mm, cauline abruptly or gradually reduced distally, margins entire (apices rounded), faces sparsely hirsutulous or villous to sparsely strigose or glabrate, sometimes sparsely glandular. |
basal (usually deciduous) and cauline; basal blades obovate-spatulate, 10–70 × 4–-14 mm; cauline gradually reduced distally, margins entire or with 2–3 pairs of teeth or lobes, faces hirsute to loosely strigoso-hirsute, sometimes sparsely glandular. |
Involucres | 5–8(–10) × 8–20 mm. |
3–4 × (5–)7–11 mm. |
Ray florets | 50–130; corollas blue to pink or purplish, rarely white, 7–11(–15) mm (mostly 1–2 mm wide), laminae coiling. |
75–150; corollas white, drying lilac, without abaxial midstripe, (2–)4–9.5 mm, laminae not coiling or reflexing (remaining relatively straight). |
Disc corollas | 2.4–4(–5) mm. |
1.8–2.3 mm (throats indurate and slightly inflated). |
Phyllaries | in 2–3 series (green or purplish), moderately to densely woolly-villous (hairs flattened, cross walls sometimes reddish), minutely glandular at least apically. |
in 3–4 series, hirsute, minutely glandular. |
Heads | 1. |
(in early season, 1–)5–100+ (buds nodding). |
Cypselae | 1.8–2.4 mm, 2-nerved, faces strigose; pappi: outer of setae, inner of (7–)10–18(–22) bristles. |
0.9–1.2 mm, 2-nerved (nerves whitish), faces sparsely strigose; pappi: outer of setae or scales, inner of 6–9(–12) bristles. |
2n | = 18, 27. |
= 18, 27, 36. |
Erigeron grandiflorus |
Erigeron divergens |
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Phenology | Flowering Jul–Aug(–Sep). | Flowering (Feb–)Apr–Oct. |
Habitat | Rocky sites, meadows, alpine or near timberline | Gravelly or sandy flats, riverbanks, meadows, disturbed sites, desert scrubland to grasslands, saltbush, blackbrush, sagebrush, pinyon-juniper, juniper-oak, oak, and ponderosa pine woodlands |
Elevation | 2900–4200 m (9500–13800 ft) | 60–2000(–2700) m (200–6600(–8900) ft) |
Distribution |
AZ; CO; ID; MT; NM; OR; UT; WY; AB; BC
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AZ; CA; CO; ID; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; Mexico (Baja California, Chihuahua, Coahuila, Durango, Sonora, and southward)
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Discussion | S. A. Spongberg (1971) recognized only the triploid populations as Erigeron grandiflorus and assigned the diploid ones to E. simplex. He hypothesized that the triploids incorporate genomic elements from an ancestor other than E. simplex. Based on his comments and annotations, however, triploids in southern Canada and the western United States apparently differ from the much more widespread diploids only quantitatively, having involucres and florets at the higher end of size ranges. Morphologic distinctions between the ploidal races do not provide a basis for consistent distinction. Spongberg (p. 200) also noted that “because of the intergrading of morphologic features of plants of Erigeron grandiflorus...the single most important criterion indicative of this taxon is highly irregular [in shape] and greatly abortive pollen.” These pollen features result from meiotic anomalies associated with the triploid condition. Specimen citations by A. Cronquist (1947) for Erigeron grandiflorus were mostly from collections of the species treated here as E. porsildii. He also cited two collections from southwestern Alberta; those and the type collection of E. grandiflorus (from the same region) are disjunct by more than 1500 kilometers from the more northern range of E. porsildii and instead lie at the northern extremity of the range of what previously has generally been identified as E. simplex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Polyploidy and agamospermy apparently are common in Erigeron divergens and contribute to the variability and, probably to some extent, the polymorphism characteristic of this species. Diploids appear to be scattered through the range of the species, at least in its southern part. The form of Erigeron divergens with persistent, deeply pinnatifid basal leaves with relatively long petioles and tendency toward perenniality (woody taproots) has been named E. accedens [occurs at 200–1000(–1200) m and flowers Feb–Apr(–May)]. Such plants have sometimes been identified as E. lobatus, and it is possible that they may have originated as hybrids with it; in most ways they are similar to E. divergens and apparently intergrade with it. The E. accedens form occurs in California, Arizona, New Mexico, southwest Texas, Baja California, northern Sonora, Durango, and Chihuahua, apparently sympatrically with more typical E. divergens, suggesting that the former is stabilized and self-reproductive (probably agamospermic). Available chromosome counts are triploid. Rare plants (e.g., E. incomptus) may have filiform rays barely extending past the involucre. Another common form of Erigeron divergens in the broad sense (mostly in Chihuahua, also southern Texas, New Mexico, and Arizona) has bowl-shaped plants with decumbent stems, a discrete taproot, persistent, lobed basal leaves, and solitary heads topping usually simple stems, and usually is early-season in phenology. Further comments related to variation in E. divergens are found under 163. Erigeron multiceps. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 324. | FNA vol. 20, p. 339. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | E. simplex | E. divaricatus, E. accedens, E. incomptus, E. solisaltator |
Name authority | Hooker: Fl. Bor.-Amer. 2: 18, plate 123. (1834) | Torrey & A. Gray: Fl. N. Amer. 2: 175. (1841) |
Web links |