Cynara cardunculus subsp. flavescens |
Cynara cardunculus |
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cardoon |
artichoke, artichoke thistle, cardoon |
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Taproots | fleshy. |
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Stems | glabrous to densely arachnoid-tomentose. |
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Leaves | armed with spines 1–3 cm. |
basal blades 30–200 cm, margins deeply 1–2-pinnately lobed or divided to nearly compound, lobes oblong to lanceolate, entire to coarsely toothed, teeth and lobes innocuous to prominently spine-tipped, spines 1–30 mm, often clustered along petiole and at base of lobes, abaxial faces densely gray- or white-tomentose, adaxial faces thinly cobwebby-tomentose; cauline leaves often short-decurrent as spiny wings. |
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Involucres | often purplish tinged, 30–150 × 40–150 mm excluding spreading phyllary tips, constricted distally or not. |
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Corollas | blue or purple (rarely white), 3–5 cm; styles long-exserted. |
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Phyllaries | lanceolate to broadly ovate, bases appressed, spreading apices obtuse to acute or acuminate, spineless or tipped with spines 1–9 mm or truncate, abruptly mucronate, and spineless or minutely spine-tipped. |
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Cypselae | 4–8 mm; pappus bristles 2–4 cm. |
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Middle | phyllaries acute to short-acuminate at apex with point 10–21 mm and spine tip 2–5(–6) mm, distal margins with prominent yellowish margins 0.5–1+ mm. |
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Cynara cardunculus subsp. flavescens |
Cynara cardunculus |
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Phenology | Flowering spring–summer (Apr–Jul). | |||||
Habitat | Disturbed areas in grasslands, coastal scrub, chaparral, riparian habitats, fallow fields, roadsides | |||||
Elevation | 0–500 m (0–1600 ft) | |||||
Distribution |
CA; w Mediterranean region; Macaronesia [Introduced in North America] |
CA; Mediterranean region; Macaronesia [Introduced in North America]
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Discussion | Artichoke thistle is a noxious weed that tends to spread aggressively in overgrazed range lands and may invade undisturbed natural vegetation as well. Dense infestations form pure stands, sometimes several hec-tares in extent. The long, needlelike spines deter herbivores and the large rosettes suppress the growth of other plants. Vigorous root sprouts can develop from fragments of the deep taproots left in the soil by cultivation or mechanical clearing. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Cynara cardunculus is a species of considerable economic importance. The globe artichoke, formerly treated as C. scolymus, was included as a horticulturally derived form of C. cardunculus (A. Wiklund 1992). The artichoke and the cardoon, another horticultural race of C. cardunculus, have been cultivated for centuries–the former for edible phyllary bases and receptacles, and the latter for edible stems and leaf rachises. That species has a darker side, however. Wild type races (artichoke thistles) are invasive and tenacious weeds that have infested Mediterranean climate areas of California, South America, South Africa, and Australia. Wiklund recognized two subspecies of C. cardunculus: subsp. cardunculus includes the artichoke, cardoon, and various wild types; subsp. flavescens includes some of the most invasive weedy members of the species. It is not certain that all of the weedy artichoke thistles in California are members of the latter subspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 90. | FNA vol. 19, p. 89. | ||||
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Name authority | Wiklund: Bot. J. Linn. Soc. 109: 120, fig. 15A–D, F–J. (1992) | Linnaeus: Sp. Pl. 2: 827. (1753) | ||||
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