Castilleja oresbia |
Castilleja elata |
|
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pale paintbrush, pale Wallowa Indian paintbrush, pale Wallowa paintbrush |
Siskiyou Indian paintbrush, Siskiyou paintbrush, slender paintbrush |
|
Habit | Herbs, perennial, 0.9–3 dm; from a woody caudex; with a stout taproot. | Herbs, perennial, 1.9–6.2 dm; from a remote woody caudex; with a taproot. |
Stems | few to several, erect or ascending, sometimes decumbent at base, unbranched or branched, hairs usually retrorse, medium length, ± soft, eglandular, mixed with very short-glandular ones, sometimes with spreading, long, soft ones. |
solitary, sometimes few, proximally creeping, becoming rhizomatous, distally ascending to erect, unbranched or branched, glabrous proximally, glabrate distally, hairs ascending, medium length, soft, eglandular. |
Leaves | green to purple, linear to lanceolate, 2–7 cm, not fleshy, margins plane, involute, 3–5(–7)-lobed, apex acuminate to acute; lobes spreading, linear to sometimes narrowly lanceolate, apex acute. |
widely spaced on stem, green, linear to lanceolate or narrowly oblong, 1.3–7.2 cm, not fleshy, margins plane, ± revolute, 0-lobed, apex broadly acute to acuminate, distalmost sometimes obtuse. |
Inflorescences | 2.5–18 × 1–3.5 cm; bracts pale green to yellow-green or pale, dull reddish brown throughout, or proximally so colored but changing gradually to cream or yellowish on distal margins, narrowly to broadly lanceolate, (3–)5–7(–9)-lobed; lobes ascending, linear, long, proximal lobes arising below mid length, central lobe apex obtuse, others acute. |
2–13.5 × 1.5–4.5 cm; bracts proximally greenish, rarely pink, distally pink, magenta, or purple, sometimes distally buff, dull yellow, cream, or light yellow-orange, ovate to elliptic or lanceolate, 0–5-lobed, sometimes also with 3 small teeth on center lobe; lobes ascending or ± spreading, narrowly lanceolate to oblong, medium length, arising near or above mid length, apex acuminate to acute, sometimes obtuse. |
Corollas | straight, 21–36 mm; tube 16–20 mm; teeth of abaxial lip often exserted, beak exserted; beak adaxially green, 4.2–5.5 mm; abaxial lip green to purple, distally white, conspicuous, slightly but noticeably pouched, often visible through front cleft, 3–5 mm, 67–100% as long as beak, puberulent; teeth erect, white, 1.8–2.1 mm. |
straight, 15–25 mm; tube 7–12 mm; beak and abaxial lip exserted above calyx lobes or ± pendently exserted from abaxial calyx cleft; beak adaxially green to yellowish, 6–11 mm; abaxial lip white to green, reduced, inconspicuous, pouches 3, small, 0.5–1.5 mm, 10–15% as long as beak; teeth ascending, white to deep green, 0.5–1 mm. |
Calyces | colored as bracts, 10–25 mm; abaxial and adaxial clefts 6–7 mm, 30–60% of calyx length, deeper than laterals, lateral 5–10 mm, 40–50% of calyx length; lobes linear, apex acute. |
whitish, green, or pink, pale colored ones tending to age pink, lobes as in bracts, 9–17 mm; abaxial and adaxial clefts 4.5–8 mm, 50% of calyx length, deeper than laterals, lateral 2–3 mm, 17–22% of calyx length; lobes linear to oblanceolate, apex acute. |
2n | = 24. |
|
Castilleja oresbia |
Castilleja elata |
|
Phenology | Flowering May–Aug. | Flowering May–Aug. |
Habitat | Dry slopes and plains, sagebrush meadows, grasslands, openings in conifer forests. | Serpentine bogs and wetlands. |
Elevation | 900–2200 m. (3000–7200 ft.) | 50–1900 m. (200–6200 ft.) |
Distribution |
ID; OR
|
CA; OR
|
Discussion | Castilleja oresbia is endemic to eastern Oregon and adjacent Idaho. It is easily confused with both varieties of C. pallescens, which also occur in sagebrush habitats. Castilleja oresbia has longer calyx lobes and softer pubescence than C. pallescens var. pallescens, although some transitional specimens are found. Castilleja oresbia has a combination of longer calyx lobes, longer pubescence, and obscurely nerved bracts, which usually serve to separate it from C. pallescens var. inverta. All three have different, though somewhat overlapping, ranges. Castilleja oresbia occasionally hybridizes with C. peckiana in Grant County, Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja elata is endemic to the Siskiyou Mountains of northwestern California and adjacent Oregon. Although often treated as a subspecies of C. miniata, it differs from that species in its shorter corolla beaks and distinctive bimodal coloration, with some populations exclusively pale yellow to pale orange and others pink-purple to magenta, as well as its specialized habitat in serpentine wetlands, where it often grows alongside Darlingtonia californica. Castilleja miniata grows on more mesic to moderately xeric substrates in the general vicinity of C. elata but with no sign of intergradation between the two species. The origin and significance of the two discrete color forms of C. elata deserve further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 635. | FNA vol. 17, p. 604. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. miniata subsp. elata | |
Name authority | Greenman: Bot. Gaz. 48: 147. (1909) | Piper: Smithsonian Misc. Collect. 50: 201. (1907) |
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