Castilleja oresbia |
Castilleja chrymactis |
|
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pale paintbrush, pale Wallowa Indian paintbrush, pale Wallowa paintbrush |
green Indian paintbrush, green paintbrush |
|
Habit | Herbs, perennial, 0.9–3 dm; from a woody caudex; with a stout taproot. | Herbs, perennial, 2.8–6.6 dm; from a woody caudex; with a taproot. |
Stems | few to several, erect or ascending, sometimes decumbent at base, unbranched or branched, hairs usually retrorse, medium length, ± soft, eglandular, mixed with very short-glandular ones, sometimes with spreading, long, soft ones. |
several to many, erect to ascending, unbranched or branched, sometimes with short, leafy axillary shoots, glabrous or hairs sparse proximally, distally spreading-erect, long, soft, eglandular, mixed with shorter ones. |
Leaves | green to purple, linear to lanceolate, 2–7 cm, not fleshy, margins plane, involute, 3–5(–7)-lobed, apex acuminate to acute; lobes spreading, linear to sometimes narrowly lanceolate, apex acute. |
green or purple-tinged, sometimes deep purple, linear-lanceolate to lanceolate or ovate, (1.5–)5–11.7 cm, not fleshy, margins plane, sometimes ± wavy, flat or involute, 0–3(–5)-lobed, apex acute to obtuse, sometimes rounded; lobes erect or ascending, linear, apex acute or acuminate. |
Inflorescences | 2.5–18 × 1–3.5 cm; bracts pale green to yellow-green or pale, dull reddish brown throughout, or proximally so colored but changing gradually to cream or yellowish on distal margins, narrowly to broadly lanceolate, (3–)5–7(–9)-lobed; lobes ascending, linear, long, proximal lobes arising below mid length, central lobe apex obtuse, others acute. |
3.5–18 × 3–7.5 cm; bracts proximally pale green to dull purplish, distally bright red or scarlet to yellow, orange, yellow-orange, and salmon, sometimes tricolored with central band of deeper purple or deeper shade of the far-distal color, broadly lanceolate, oblong, elliptic, or oblong, 0–3(–9)-lobed; lobes ascending, lanceolate, triangular, or oblong, short, arising at or above mid length, central lobe apex rounded, lateral ones acute to obtuse. |
Corollas | straight, 21–36 mm; tube 16–20 mm; teeth of abaxial lip often exserted, beak exserted; beak adaxially green, 4.2–5.5 mm; abaxial lip green to purple, distally white, conspicuous, slightly but noticeably pouched, often visible through front cleft, 3–5 mm, 67–100% as long as beak, puberulent; teeth erect, white, 1.8–2.1 mm. |
straight to slightly curved, 26–43 mm; tube 16–24 mm; beak exserted, adaxially green to yellow, 9–19 mm; abaxial lip whitish or dark green, reduced, not exserted, sometimes visible through front calyx cleft, 1.5–2 mm, 20% as long as beak, glabrous or sparsely hairy, hairs spreading; teeth reduced, incurved, deep green, 0.5–1 mm. |
Calyces | colored as bracts, 10–25 mm; abaxial and adaxial clefts 6–7 mm, 30–60% of calyx length, deeper than laterals, lateral 5–10 mm, 40–50% of calyx length; lobes linear, apex acute. |
colored as bracts, sometimes more deeply pigmented and contrasting with bract coloration, 24–31 mm; abaxial and adaxial clefts 12–19 mm, 45–65% of calyx length, deeper than laterals, lateral 7–11 mm, 25–32% of calyx length; lobes lanceolate, narrowly triangular, or almost oblong, apex acute, sometimes obtuse or acuminate. |
Castilleja oresbia |
Castilleja chrymactis |
|
Phenology | Flowering May–Aug. | Flowering (Jan–)Feb–Sep(–Oct). |
Habitat | Dry slopes and plains, sagebrush meadows, grasslands, openings in conifer forests. | Glacial outwash plains, river flats, moist or wet openings, beach meadows, roadsides. |
Elevation | 900–2200 m. (3000–7200 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
ID; OR
|
AK |
Discussion | Castilleja oresbia is endemic to eastern Oregon and adjacent Idaho. It is easily confused with both varieties of C. pallescens, which also occur in sagebrush habitats. Castilleja oresbia has longer calyx lobes and softer pubescence than C. pallescens var. pallescens, although some transitional specimens are found. Castilleja oresbia has a combination of longer calyx lobes, longer pubescence, and obscurely nerved bracts, which usually serve to separate it from C. pallescens var. inverta. All three have different, though somewhat overlapping, ranges. Castilleja oresbia occasionally hybridizes with C. peckiana in Grant County, Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja chrymactis is restricted to the coast of southeastern Alaska, from Glacier Bay to Yakutat. In some respects, it resembles a hybrid swarm between C. miniata and C. unalaschcensis. If C. chrymactis originated as a hybrid, at present it has formed a number of populations and has a stable morphology that differs from both putative parents, especially in calyx structure. Reports of its introduction to Point Reyes, California (T. I. Chuang and L. R. Heckard 1993b), are based on C. leschkeana, a distinctive species known only from two records near the type locality and presumed to be extinct. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 635. | FNA vol. 17, p. 596. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Greenman: Bot. Gaz. 48: 147. (1909) | Pennell: Proc. Acad. Nat. Sci. Philadelphia 86: 539. (1934) |
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