Castilleja elegans |
Castilleja crista-galli |
|
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elegant Indian paintbrush, elegant paintbrush |
cock's-comb paintbrush, mountainside Indian paintbrush |
|
Habit | Herbs, perennial, 0.5–3 dm; from a woody caudex; with a slender taproot. | Herbs, perennial, 1–5 dm; from a woody caudex; with a taproot. |
Stems | few to several, ascending to erect, or slightly decumbent at base, unbranched, hairy, sometimes glabrate near base, hairs erect-ascending, whitish or yellowish, long, soft, eglandular, mixed with shorter stipitate-glandular ones, especially distally. |
few to several, erect or ascending, unbranched or branched, sometimes with short, leafy axillary shoots, hairy, sometimes glabrate proximally, hairs spreading to retrorse, medium length to long, soft, eglandular, often mixed distally with shorter stipitate-glandular ones. |
Leaves | green to sometimes purple-tinged, lanceolate to narrowly oblong, (1.2–)2–6(–9) cm, not fleshy, margins plane, sometimes ± wavy, flat to ± involute, 0-lobed, sometimes 3-lobed distally, immediately below inflorescence, apex acuminate to caudate; lobes ascending to erect, linear to lanceolate, lateral lobes narrower than central, apex acuminate. |
green, linear to narrowly lanceolate, 2–8 cm, not fleshy, margins plane, involute, 0–5-lobed, apex acute; lateral lobes spreading, linear, apex acuminate. |
Inflorescences | 1.5–9 × 1.5–3 cm; bracts purple to pink-purple or reddish purple throughout, sometimes proximally purple to pink-purple or reddish purple, distally whitish or pale pink, oblong to lanceolate or narrowly ovate, (0–)3–7-lobed; lobes spreading-ascending, linear-lanceolate, narrow, short or long, arising near or above mid length, center lobe apex rounded to sometimes acute, lateral ones acute. |
3–6(–11) × 1.5–6.5 cm; bracts proximally greenish, distally red, red-orange, or orange, sometimes yellow or dull salmon, narrowly to broadly lanceolate or oblong, 3–5-lobed; lobes ascending-spreading, linear-lanceolate, long, arising below mid length, central lobe apex rounded to obtuse, lateral ones acute. |
Corollas | straight, 18–28 mm; tube 17–20 mm; abaxial lip generally visible between calyx lobes, sometimes exserted above them, beak exserted; beak adaxially green, yellowish green, or purplish green, 4–8 mm; abaxial lip purple or magenta, medium sized, often visible through or above abaxial cleft, pouches 3, slightly inflated, 3–5.5 mm, 50+% as long as beak; teeth ascending, purple or magenta, 1–1.5 mm. |
straight, (25–)30–40(–45) mm; tube 15–20 mm; abaxial lip visible through front cleft, beak long-exserted from calyx; beak adaxially green or yellow-green, 16–21 mm; abaxial lip proximally white or yellow-green, distally green, reduced, usually visible in front cleft, 3 mm, 20% as long as beak; teeth incurved to ascending, green, 1 mm. |
Calyces | yellow to yellow-green, rarely purplish to dull red throughout, 19–25 mm; abaxial and adaxial clefts 10–17 mm, 60–70% of calyx length, deeper than laterals, lateral (2–)5–8 mm, 20–25% of calyx length; lobes narrowly oblong to lanceolate, apex rounded or obtuse, sometimes acute. |
colored as bracts, (20–)25–35 mm; abaxial and adaxial clefts (6–)10–17 mm, 50% of calyx length, deeper than laterals, lateral (1–)3–6(–10) mm, 35% of calyx length; lobes slender, triangular, apex acute. |
2n | = 24. |
= 96. |
Castilleja elegans |
Castilleja crista-galli |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Meadows, tundra, fell fields, moraines, talus, rocky slopes, shrub thickets, gravel bars, lakeshores. | Rocky slopes, talus, ridges, dry to moist, open, conifer forests, montane meadows. |
Elevation | 0–2000 m. (0–6600 ft.) | 1500–2900 m. (4900–9500 ft.) |
Distribution |
AK; MB; NT; NU; QC; YT; e Asia (Russian Far East) |
ID; MT; WY
|
Discussion | Castilleja elegans is a characteristic species of the arctic and arctic-alpine regions. Its eastern and western limits are poorly understood, due to confusion with the similar species C. raupii to the east and south, and with C. rubra (Drobow) Rebristaya in eastern Asia. All three are part of a morphologically variable complex of recently evolved and poorly differentiated entities informally known as the C. pallida complex, after the first-described member of the group. Some reports of C. elegans in eastern Canada are here referred to C. septentrionalis. A recent collection of C. elegans from the northern Canadian Rocky Mountains is apparently disjunct, though it may also suggest its wider presence in Alberta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Castilleja crista-galli is found in the Rocky Mountains of southwestern Montana and northwestern Wyoming. The extent of its distribution into adjacent Idaho is unresolved, in part because it is frequently confused with either C. linariifolia or C. miniata. Castilleja crista-galli appears to be morphologically intermediate between them, leading to speculation that it might be an allopolyploid derivative. A DNA study (S. Matthews and M. Lavin 1998) showed little support for a hybrid origin. Castilleja crista-galli may be separated with some difficulty from the other two species by the presence of at least some short hairs on the stems and the frequently three- to five-parted leaves. Castilleja linariifolia and C. miniata both usually have subglabrous stems and entire leaves, sometimes three-parted distally, near the inflorescence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 604. | FNA vol. 17, p. 599. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. pallida subsp. elegans | |
Name authority | Malte: Rhodora 36: 187. (1934) | Rydberg: Mem. New York Bot. Gard. 1: 355. (1900) |
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