Cardamine diphylla |
Cardamine flagellifera |
|
---|---|---|
broadleaf toothwort, crinkleroot, two-leaf toothwort |
Blue Ridge bittercress |
|
Habit | Perennials; usually glabrous, rarely sparsely pubescent. | Perennials; hirsute or pilose proximally, sparsely pubescent or glabrous distally. |
Rhizomes | (unsegmented), cylindrical, 2–10 mm diam., (somewhat uniform, fleshy, not fragile, with dentate leaf scars). |
swollen, compact at stem base, (fleshy, stolons many, slender, ca. 1 mm diam., pubescent or glabrous). |
Stems | erect, unbranched, (1.2–)1.5–3.5(–4) dm, rarely sparsely pubescent distally. |
erect, usually unbranched, rarely branched distally, 1–2.5 dm, sparsely to densely hirsute or pilose proximally, sparsely so or glabrous distally. |
Rhizomal leaves | 3-foliolate, (5.5–)8–22(–26) cm, leaflets petiolulate or subsessile; petiole (3–)4.5–16(–20) cm; lateral leaflets subsessile or petiolulate, blade often similar to terminal, base sometimes oblique; terminal leaflet (petiolule 0.5–1.2 cm), blade ovate-elliptic to broadly ovate, (2–)3.5–8(–10) cm × (5–)20–65(–80) mm, base cuneate to obtuse, margins coarsely crenate or dentate, (surfaces puberulent, trichomes to 0.1 mm). |
usually simple, rarely 3-foliolate, (3–)6–16 cm, leaflets petiolulate or subsessile; petiole (1.5–)4–13 cm; lateral leaflets subsessile or petiolulate (to 0.5 cm), blade similar to terminal, much smaller; terminal leaflet (petiolule 0.5–2 cm), blade orbicular to reniform or broadly ovate, (1–)1.5–4.5 cm, base cordate, margins repand to coarsely crenate, (apiculate at vein endings, surfaces sparsely pubescent or glabrous). |
Cauline leaves | 2 (or 3) [(sub)opposite], 3-foliolate, (similar to rhizomal leaves), petiolate, leaflets petiolulate or subsessile; petiole (1–)2–4.5 cm, base not auriculate; lateral leaflets similar to terminal; terminal leaflet subsessile or petiolulate (0.2–1 cm), blade broadly elliptic to ovate, (2–)4–8(–10) cm × 10–50 mm, margins coarsely dentate or crenate, (margins minutely puberulent). |
2–5, 3 or 5-foliolate, petiolate, leaflets petiolulate or subsessile; petiole 0.8–5 cm, base not auriculate; lateral leaflets sessile or petiolulate (to 0.5 cm), blade similar to terminal, smaller; terminal leaflet petiolulate (0.5–1.5 cm), blade broadly ovate to suborbicular, 1.5–4(–5) cm × 12–35 mm, margins repand to coarsely crenate or slightly lobed, (apiculate at vein endings, margins glabrous). |
Racemes | ebracteate. |
ebracteate. |
Flowers | sepals oblong, (4–)5–8 × 2–3 mm, lateral pair slightly saccate basally; petals white or pink to purple, obovate to oblanceolate, (7–)9–15(–17) × (3–)4–7 mm, (short-clawed, apex rounded); filaments: median pairs 5–8 mm, lateral pair 3.5–6 mm; anthers linear to oblong, 2.5–3 mm. |
sepals (ascending) oblong, 3–3.5 × 1–1.5 mm, lateral pair not saccate basally; petals white, oblanceolate, 5–6.5(–8) × 1.5–2.5 mm, (not clawed, apex obtuse to subemarginate); filaments: median pairs 3.5–5 mm, lateral pair 2.5–3 mm; anthers oblong, 1.2–1.6 mm. |
Fruiting pedicels | ascending to divaricate, (10–)15–30(–36) mm. |
ascending to divaricate-ascending, 7–20 mm. |
Fruits | linear-lanceolate, 1.5–4 cm × 1.5–2.5 mm; ovules 10–14 per ovary; style 4–8(–10) mm. |
linear, 1.5–2.5 cm × 1–1.2 mm; ovules 10–14 per ovary; style 1.2–2.5 mm. |
Seeds | (rarely produced) brown, oblong, 2–2.2 × 1.2–1.5 mm (cotyledons incumbent). |
brown, oblong, 1.2–1.7 × 0.8–1.1 mm. |
2n | = 96. |
|
Cardamine diphylla |
Cardamine flagellifera |
|
Phenology | Flowering Apr–Jun. | Flowering Mar–Jun. |
Habitat | Wooded bottoms and ravines, cliffs, bluffs, ledges, shaded slopes, meadows, moist fields, alluvial banks, rich woods | Moist wooded slopes, gorges, wooded ravines, seepage places |
Elevation | 50-1300 m (200-4300 ft) | 300-1000 m (1000-3300 ft) |
Distribution |
AL; AR; CT; GA; IL; IN; KY; MA; ME; MI; MN; NC; NH; NJ; NY; OH; PA; SC; TN; VA; VT; WI; WV; NB; NS; ON; QC
|
GA; NC; SC; TN; WV |
Source | FNA vol. 7, p. 472. | FNA vol. 7, p. 474. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Dentaria diphylla, Dentaria bifolia, Dentaria incisa | C. flagellifera var. hugeri, C. hugeri |
Name authority | (Michaux) Alph. Wood: Amer. Bot. Fl., 37. (1870) | O. E. Schulz: Bot. Jahrb. Syst. 32: 405. (1903) |
Web links |