Cardamine diphylla |
Brassicaceae tribe Cardamineae |
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broadleaf toothwort, crinkleroot, two-leaf toothwort |
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Habit | Perennials; usually glabrous, rarely sparsely pubescent. | Annuals, biennials, or perennials; eglandular. |
Rhizomes | (unsegmented), cylindrical, 2–10 mm diam., (somewhat uniform, fleshy, not fragile, with dentate leaf scars). |
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Stems | erect, unbranched, (1.2–)1.5–3.5(–4) dm, rarely sparsely pubescent distally. |
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Rhizomal leaves | 3-foliolate, (5.5–)8–22(–26) cm, leaflets petiolulate or subsessile; petiole (3–)4.5–16(–20) cm; lateral leaflets subsessile or petiolulate, blade often similar to terminal, base sometimes oblique; terminal leaflet (petiolule 0.5–1.2 cm), blade ovate-elliptic to broadly ovate, (2–)3.5–8(–10) cm × (5–)20–65(–80) mm, base cuneate to obtuse, margins coarsely crenate or dentate, (surfaces puberulent, trichomes to 0.1 mm). |
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Cauline leaves | 2 (or 3) [(sub)opposite], 3-foliolate, (similar to rhizomal leaves), petiolate, leaflets petiolulate or subsessile; petiole (1–)2–4.5 cm, base not auriculate; lateral leaflets similar to terminal; terminal leaflet subsessile or petiolulate (0.2–1 cm), blade broadly elliptic to ovate, (2–)4–8(–10) cm × 10–50 mm, margins coarsely dentate or crenate, (margins minutely puberulent). |
(rarely absent), usually petiolate, sometimes sessile; blade (simple or compound), base auriculate or not, margins entire or dentate to pinnately lobed. |
Trichomes | absent or simple. |
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Racemes | ebracteate. |
usually ebracteate (Selenia bracteate throughout), often elongated in fruit. |
Flowers | sepals oblong, (4–)5–8 × 2–3 mm, lateral pair slightly saccate basally; petals white or pink to purple, obovate to oblanceolate, (7–)9–15(–17) × (3–)4–7 mm, (short-clawed, apex rounded); filaments: median pairs 5–8 mm, lateral pair 3.5–6 mm; anthers linear to oblong, 2.5–3 mm. |
actinomorphic; sepals erect to spreading or ascending, lateral pair seldom saccate basally; petals white, yellow, pink, lilac, or purple, claw usually present, rarely absent, often distinct; filaments unappendaged, not winged; pollen 3-colpate. |
Fruiting pedicels | ascending to divaricate, (10–)15–30(–36) mm. |
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Fruits | linear-lanceolate, 1.5–4 cm × 1.5–2.5 mm; ovules 10–14 per ovary; style 4–8(–10) mm. |
silicles or siliques, dehiscent, unsegmented, terete, 4-angled, or latiseptate, rarely angustiseptate; ovules 4–300[–numerous] per ovary; style distinct or obsolete (absent in Subularia); stigma usually entire. |
Seeds | (rarely produced) brown, oblong, 2–2.2 × 1.2–1.5 mm (cotyledons incumbent). |
biseriate or uniseriate; cotyledons usually accumbent, sometimes incumbent. |
2n | = 96. |
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Cardamine diphylla |
Brassicaceae tribe Cardamineae |
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Phenology | Flowering Apr–Jun. | |
Habitat | Wooded bottoms and ravines, cliffs, bluffs, ledges, shaded slopes, meadows, moist fields, alluvial banks, rich woods | |
Elevation | 50-1300 m (200-4300 ft) | |
Distribution |
AL; AR; CT; GA; IL; IN; KY; MA; ME; MI; MN; NC; NH; NJ; NY; OH; PA; SC; TN; VA; VT; WI; WV; NB; NS; ON; QC
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Nearly worldwide |
Discussion | Genera 12, species ca. 335 (10 genera, 85 species in the flora). The assignment of Subularia to Cardamineae is provisional and based solely on morphology. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 472. | FNA vol. 7, p. 458. |
Parent taxa | ||
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Dentaria diphylla, Dentaria bifolia, Dentaria incisa | |
Name authority | (Michaux) Alph. Wood: Amer. Bot. Fl., 37. (1870) | Dumortier: Fl. Belg., 124. (1827) |
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