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broadleaf toothwort, crinkleroot, two-leaf toothwort

Habit Perennials; usually glabrous, rarely sparsely pubescent.
Rhizomes

(unsegmented), cylindrical, 2–10 mm diam., (somewhat uniform, fleshy, not fragile, with dentate leaf scars).

Stems

erect, unbranched, (1.2–)1.5–3.5(–4) dm, rarely sparsely pubescent distally.

Rhizomal leaves

3-foliolate, (5.5–)8–22(–26) cm, leaflets petiolulate or subsessile;

petiole (3–)4.5–16(–20) cm;

lateral leaflets subsessile or petiolulate, blade often similar to terminal, base sometimes oblique;

terminal leaflet (petiolule 0.5–1.2 cm), blade ovate-elliptic to broadly ovate, (2–)3.5–8(–10) cm × (5–)20–65(–80) mm, base cuneate to obtuse, margins coarsely crenate or dentate, (surfaces puberulent, trichomes to 0.1 mm).

Cauline leaves

2 (or 3) [(sub)opposite], 3-foliolate, (similar to rhizomal leaves), petiolate, leaflets petiolulate or subsessile;

petiole (1–)2–4.5 cm, base not auriculate;

lateral leaflets similar to terminal;

terminal leaflet subsessile or petiolulate (0.2–1 cm), blade broadly elliptic to ovate, (2–)4–8(–10) cm × 10–50 mm, margins coarsely dentate or crenate, (margins minutely puberulent).

Racemes

ebracteate.

Flowers

sepals oblong, (4–)5–8 × 2–3 mm, lateral pair slightly saccate basally;

petals white or pink to purple, obovate to oblanceolate, (7–)9–15(–17) × (3–)4–7 mm, (short-clawed, apex rounded);

filaments: median pairs 5–8 mm, lateral pair 3.5–6 mm;

anthers linear to oblong, 2.5–3 mm.

Fruiting pedicels

ascending to divaricate, (10–)15–30(–36) mm.

Fruits

linear-lanceolate, 1.5–4 cm × 1.5–2.5 mm;

ovules 10–14 per ovary;

style 4–8(–10) mm.

Seeds

(rarely produced) brown, oblong, 2–2.2 × 1.2–1.5 mm (cotyledons incumbent).

2n

= 96.

Cardamine diphylla

Phenology Flowering Apr–Jun.
Habitat Wooded bottoms and ravines, cliffs, bluffs, ledges, shaded slopes, meadows, moist fields, alluvial banks, rich woods
Elevation 50-1300 m (200-4300 ft)
Distribution
from FNA
AL; AR; CT; GA; IL; IN; KY; MA; ME; MI; MN; NC; NH; NJ; NY; OH; PA; SC; TN; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 472.
Parent taxa Brassicaceae > tribe Cardamineae > Cardamine
Sibling taxa
C. angulata, C. angustata, C. bellidifolia, C. blaisdellii, C. breweri, C. bulbosa, C. californica, C. clematitis, C. concatenata, C. constancei, C. cordifolia, C. digitata, C. dissecta, C. douglassii, C. flagellifera, C. flexuosa, C. hirsuta, C. holmgrenii, C. impatiens, C. longii, C. macrocarpa, C. maxima, C. micranthera, C. microphylla, C. nuttallii, C. nymanii, C. occidentalis, C. oligosperma, C. pachystigma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica, C. pratensis, C. purpurea, C. rotundifolia, C. rupicola, C. umbellata
Synonyms Dentaria diphylla, Dentaria bifolia, Dentaria incisa
Name authority (Michaux) Alph. Wood: Amer. Bot. Fl., 37. (1870)
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