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doll's-daisy

Habit Perennials, 30–200+ cm (rhizomatous, stoloniferous, or with basal offshoots).
Stems

erect to ascending, branched distally (ribbed), glabrous.

Leaves

basal (withering by flowering) and cauline; alternate;

sessile (sometimes decurrent);

blades 1-nerved, linear, subulate to lanceolate or oblanceolate (bases attenuate, cauline sometimes decurrent), margins entire to serrulate (teeth callous-tipped), (apices acute to acuminate) faces glabrous.

Receptacles

conic to hemispheric, obscurely pitted, epaleate.

Ray florets

20–60, pistillate, fertile;

corollas white to lilac (laminae linear to elliptic).

Disc florets

50–400+, bisexual, fertile;

corollas yellow, tubes shorter than funnelform throats, lobes 5, erect, triangular;

style-branch appendages deltate.

Phyllaries

30–55 in 3–6 series, 1-nerved (midribs thickened, resinous; flat), linear to oblanceolate or spatulate, unequal to subequal, margins ± membranaceous, faces glabrous.

Heads

radiate, (10–100+) in leafy, corymbiform or paniculiform arrays. (Peduncles bracteate or ebracteate, bracts leaflike or scalelike.) Involucres hemispheric, (2–5 ×) 3–14 mm.

Cypselae

dimorphic (brownish);

ray ± triquetrous, margins arrowly winged, ribs 2 abaxial, 1 adaxial;

disc obovoid to cuneate or obcordiform, laterally compressed, margins winged or not, resin ducts present, faces often puberulent;

pappi persistent, usually of 2–3 sclerified awns plus (0–)7–12 shorter bristles or scales in 1 series.

x

= 9.

Boltonia

Distribution
from USDA
North America
[BONAP county map]
Discussion

Species 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Disc cypselae lacking wings, or if present wings 0.01–0.1 mm wide; pappus awns 0–0.4 mm
→ 2
1. Disc cypselae wings 0.1–0.5 mm wide; pappus awns (0.3–)0.4–2 mm
→ 3
2. Heads in corymbiform arrays; outer phyllaries 1.7–2.8 mm; ray laminae 5–13 mm
B. asteroides
2. Heads in paniculiform arrays; outer phyllaries 1.2–1.6 mm; ray laminae 3–7.5 mm
B. caroliniana
3. Bases of cauline leaves decurrent; rhizomes and stolons absent; Illinois, Missouri
B. decurrens
3. Bases of cauline leaves not decurrent; rhizomes or stolons present or absent; widely distributed or not in Illinois or Missouri.
→ 4
4. Heads in paniculiform arrays, primary branches mostly spreading; only proximal bracts leaflike, 8–55 mm, others reduced and linear to linear-oblanceolate or subulate, 2–10(–15) mm
B. diffusa
4. Heads in corymbiform arrays, branches spreading to ascending; most bracts leaflike, 10–120 mm, only peduncle bracts reduced, linear to subulate or ovate to linear-oblanceolate, 1–12(–62) mm
→ 5
5. Pappus awns 0.3–0.8 mm; outer phyllaries 1.3–2 mm; ray laminae 5–9 mm; Florida panhandle to se Louisiana
B. apalachicolensis
5. Pappus awns 0.4–2 mm; outer phyllaries 1.5–3.8 mm; ray laminae 5–13 mm; e North America, Idaho, Oregon
B. asteroides
Source FNA vol. 20, p. 353. Authors: Vesna Karaman-Castro, Lowell E. Urbatsch.
Parent taxa Asteraceae > tribe Astereae
Subordinate taxa
B. apalachicolensis, B. asteroides, B. caroliniana, B. decurrens, B. diffusa
Name authority L’Héritier: Sert. Angl., 27. (1789)
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