Boechera williamsii |
Boechera rubicundula |
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Mount Day rockcress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette, often somewhat elevated above ground surface on woody base, 1–5 dm, densely pubescent proximally, trichomes simple, 0.5–1 mm, mixed with long-stalked, 2- or 3-rayed ones, 0.1–0.5 mm, moderately to sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
blade oblanceolate, 4–8 mm wide, margins denticulate, ciliate along petiole, trichomes to 1 mm, surfaces densely pubescent, trichomes long-stalked, 2–5-rayed, 0.1–0.5 mm. |
Cauline leaves | 6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
8–25, often concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves moderately pubescent. |
Racemes | 5–17-flowered, unbranched. |
12–34-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals purplish, 6–8 × 2–2.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect, straight, 2–4(–7) mm, glabrous. |
divaricate-ascending to horizontal, straight, 5–10 mm, pubescent, trichomes spreading, simple and 2-rayed. |
Fruits | erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
divaricate-ascending to horizontal, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 4–8 cm × 1.7–2.2 mm; valves glabrous; ovules 70–102 per ovary; style 0.1–0.3 mm. |
Seeds | sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
uniseriate, 1.4–1.8 × 1–1.3 mm; wing continuous, 0.1–0.2 mm wide. |
Boechera williamsii |
Boechera rubicundula |
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Phenology | Flowering May–Jun. | Flowering Apr–May. |
Habitat | Gravelly soil in sagebrush-grassland communities | Mountain slopes |
Elevation | 2300-2800 m (7500-9200 ft) | ca. 1200 m (ca. 3900 ft) |
Distribution |
WY |
CA |
Discussion | Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera rubicundula is an apomictic species that arose through hybridization between B. arcuata and B. breweri (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). It is known only from the type locality on Mt. Day in west-central California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 404. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis williamsii | Arabis arcuata var. rubicundula |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (Jepson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 273. (2007) |
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