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broad-seed rock-cress, flat-seed rock-cress, pioneer rock-cress

Habit Perennials; short-lived; sexual; caudex not woody. Perennials; long-lived; sexual; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout.

usually 1 per caudex branch, arising from center of rosette, elevated on woody base above or near ground surface, 0.6–2.8(–3.5) dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm.

blade narrowly oblanceolate, 3–7(–10) mm wide, margins entire, ciliate near petiole base, trichomes (simple or spurred), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed 0.1–0.3 mm.

Cauline leaves

6–11, sometimes concealing stem proximally;

blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous.

3–7(–12), not concealing stem;

blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent.

Racemes

5–17-flowered, unbranched.

2–7-flowered, unbranched.

Flowers

erect at anthesis;

sepals glabrous;

petals white, 3–4 × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals creamy white to purplish, 4–6 × 1–2 mm, sparsely pubescent (often some trichomes abaxially);

pollen ellipsoid.

Fruiting pedicels

erect, straight, 2–4(–7) mm, glabrous.

ascending, straight, 3–13 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

Fruits

erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm;

valves glabrous;

ovules 60–96 per ovary;

style 0.05–0.2 mm.

ascending, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), (2.5–)4–8.5 cm × 3–5.5 mm;

valves glabrous;

ovules 16–44 per ovary;

style 0.05–1 mm.

Seeds

sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally.

uniseriate, 3–6(–8) × 2–3.5 mm;

wing continuous, 0.8–2.5 mm wide.

2n

= 14.

Boechera williamsii

Boechera platysperma

Phenology Flowering May–Jun. Flowering Jun–Aug.
Habitat Gravelly soil in sagebrush-grassland communities Rock outcrops and gravelly soil in dry pine forests and lodgepole-chaparral woodlands
Elevation 2300-2800 m (7500-9200 ft) 1600-3000 m (5200-9800 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Previous authors have adopted broader circumscriptions of Boechera platysperma, including plants herein called B. covillei, B. elkoensis, and B. howellii. In our opinion, the differences between these taxa are sufficient to justify their treatment as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007, 2007b for detailed comparison). Reports of B. platysperma from Nevada (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) are based on plants of B. howellii or apomictic hybrids with other species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 411. FNA vol. 7, p. 397.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis williamsii Arabis platysperma, Arabis inamoena, Arabis oligantha
Name authority (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) (A. Gray) Al-Shehbaz: Novon 13: 388. (2003)
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