Boechera williamsii |
Boechera platysperma |
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broad-seed rock-cress, flat-seed rock-cress, pioneer rock-cress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette, elevated on woody base above or near ground surface, 0.6–2.8(–3.5) dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
blade narrowly oblanceolate, 3–7(–10) mm wide, margins entire, ciliate near petiole base, trichomes (simple or spurred), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed 0.1–0.3 mm. |
Cauline leaves | 6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
3–7(–12), not concealing stem; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 5–17-flowered, unbranched. |
2–7-flowered, unbranched. |
Flowers | erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals creamy white to purplish, 4–6 × 1–2 mm, sparsely pubescent (often some trichomes abaxially); pollen ellipsoid. |
Fruiting pedicels | erect, straight, 2–4(–7) mm, glabrous. |
ascending, straight, 3–13 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
ascending, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), (2.5–)4–8.5 cm × 3–5.5 mm; valves glabrous; ovules 16–44 per ovary; style 0.05–1 mm. |
Seeds | sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
uniseriate, 3–6(–8) × 2–3.5 mm; wing continuous, 0.8–2.5 mm wide. |
2n | = 14. |
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Boechera williamsii |
Boechera platysperma |
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Phenology | Flowering May–Jun. | Flowering Jun–Aug. |
Habitat | Gravelly soil in sagebrush-grassland communities | Rock outcrops and gravelly soil in dry pine forests and lodgepole-chaparral woodlands |
Elevation | 2300-2800 m (7500-9200 ft) | 1600-3000 m (5200-9800 ft) |
Distribution |
WY |
CA; OR
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Discussion | Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Previous authors have adopted broader circumscriptions of Boechera platysperma, including plants herein called B. covillei, B. elkoensis, and B. howellii. In our opinion, the differences between these taxa are sufficient to justify their treatment as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007, 2007b for detailed comparison). Reports of B. platysperma from Nevada (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) are based on plants of B. howellii or apomictic hybrids with other species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 397. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis williamsii | Arabis platysperma, Arabis inamoena, Arabis oligantha |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (A. Gray) Al-Shehbaz: Novon 13: 388. (2003) |
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