Boechera williamsii |
Boechera cusickii |
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Cusick's rock-cress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; (often cespitose); sexual; caudex usually woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1–2.5(–3.8) dm, densely pubescent proximally, trichomes simple, 0.5–1.5 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
blade linear to narrowly oblanceolate, 0.5–2 mm wide, margins entire, ciliate, trichomes (simple), 1–1.7 mm, surfaces moderately pubescent, trichomes simple and long-stalked, 2-rayed, 0.5–1.7 mm. |
Cauline leaves | 6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
10–30, often concealing stem proximally; blade auricles 0–0.5 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 5–17-flowered, unbranched. |
5–14-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white to lavender, 6–10 × 2–3 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect, straight, 2–4(–7) mm, glabrous. |
ascending, straight, 4–15(–20) mm, glabrous or sparsely pubescent, trichomes simple. |
Fruits | erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
ascending to divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 4–8(–9.5) cm × 2.3–3 mm; valves glabrous; ovules 56–106 per ovary; style 0.1–0.5 mm. |
Seeds | sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
uniseriate, 1.5–2.5 × 1.5–2 mm; wing continuous, 0.1–0.2 mm wide distally. |
Boechera williamsii |
Boechera cusickii |
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Phenology | Flowering May–Jun. | Flowering Apr-mid May. |
Habitat | Gravelly soil in sagebrush-grassland communities | Basaltic bluffs, rocky slopes, rock crevices, gravelly hillsides, sagebrush hills, outcrops of volcanic rock |
Elevation | 2300-2800 m (7500-9200 ft) | 600-1800 m (2000-5900 ft) |
Distribution |
WY |
ID; NV; OR; WA
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Discussion | Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Peripheral populations of Boechera cusickii in south-central Idaho and northern Nevada have a higher proportion of branched hairs, possibly resulting from hybridization with B. sparsiflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 372. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis williamsii | Arabis cusickii |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (S. Watson) Al-Shehbaz: Novon 13: 384. (2003) |
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