Boechera williamsii |
Boechera crandallii |
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Crandall's rockcress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; (often cespitose); sexual; caudex somewhat woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
usually 2–5 per caudex branch, arising from margin of rosette near ground surface, (1–)1.5–4 dm, densely pubescent proximally, trichomes short-stalked, 5–8-rayed, 0.1–0.2 mm, moderately to sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
blade narrowly oblanceolate, 1.5–3(–5) mm wide, margins entire, ciliate along petiole base, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 5–8-rayed 0.1–0.2 mm. |
Cauline leaves | 6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
5–14, not concealing stem; blade auricles 0.1–0.5 mm, surfaces of distalmost leaves pubescent. |
Racemes | 5–17-flowered, unbranched. |
10–30-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals usually white, 5–7 × 1–2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect, straight, 2–4(–7) mm, glabrous. |
ascending to divaricate-ascending, straight, 5–10 mm, pubescent, trichomes branched. |
Fruits | erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 3–5.5 cm × 0.9–1.2 mm; valves glabrous; ovules 56–84 per ovary; style 0.1–0.5 mm. |
Seeds | sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
uniseriate, 0.8–1 × 0.6–0.9 mm; wing continuous (rarely absent), to 0.1 mm wide. |
2n | = 14. |
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Boechera williamsii |
Boechera crandallii |
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Phenology | Flowering May–Jun. | Flowering May–Jun. |
Habitat | Gravelly soil in sagebrush-grassland communities | Rocky slopes and gravelly soil in sagebrush, mountain shrub, open conifer forests |
Elevation | 2300-2800 m (7500-9200 ft) | 2000-2700 m (6600-8900 ft) |
Distribution |
WY |
CO
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Discussion | Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Boechera crandallii is a sexual diploid known only from the Gunnison Basin of west-central Colorado. Despite its narrow range, the species appears to hybridize frequently, most notably with B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 372. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis williamsii | Arabis crandallii, Arabis stenoloba |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (B. L. Robinson) W. A. Weber: Phytologia 51: 369. (1982) |
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