Boechera ultra-alsa |
Boechera pinetorum |
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Holboell's rock-cress, woodland rock-cress |
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Habit | Perennials; long-lived; reproductive mode unknown; caudex woody. | Perennials; short-lived; apomictic; caudex not woody. |
Stems | 1 per caudex branch, arising from center of rosette near ground surface, ca. 1 dm, densely pubescent proximally, trichomes stalked, (2 or) 3–6-rayed, 0.2–0.5 mm, glabrate distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–8(–9.6) dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade oblanceolate, 4–6 mm wide, margins entire, not ciliate, surfaces moderately pubescent, trichomes long-stalked, 3–6 (or 7)-rayed, 0.2–0.5 mm. |
blade oblanceolate, 2–8(–11) mm wide, margins entire or denticulate, ciliate proximally, trichomes (simple), 0.5–1 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed, 0.2–0.5 mm, these mixed with fewer simple ones. |
Cauline leaves | 2–5, concealing stem proximally; blade auricles absent, surfaces of distalmost leaves pubescent. |
7–33, often concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
Racemes | 3- or 4-flowered, unbranched. |
15–63-flowered, usually unbranched. |
Flowers | not seen. |
divaricate to pendent at anthesis; sepals pubescent; petals white to lavender, 5–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect-ascending, straight, 4–5 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
reflexed, abruptly recurved near base, otherwise straight, 4–12 mm, sparsely pubescent or glabrous, trichomes spreading, simple and 2-rayed. |
Fruits | erect-ascending, sometimes appressed to rachis, not secund, straight, edges parallel, 3–4 cm × ca. 5 mm; valves glabrous; ovules ca. 16 per ovary; style 0.5–0.7 mm. |
reflexed to closely pendent, not appressed to rachis, not secund, straight or curved, edges parallel, (4.5–)5.5–8.5 cm × 1.5–2 mm; valves glabrous; ovules 70–110 per ovary; style 0.05–0.4 mm. |
Seeds | uniseriate, 5.5–6.5 × 4–4.5 mm; wing continuous, 1–2 mm wide. |
uniseriate, 1.5–1.8 × 1.2–1.5 mm; wing continuous, 0.15–0.3 mm wide. |
Boechera ultra-alsa |
Boechera pinetorum |
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Phenology | Flowering Jun–Jul. | Flowering May–Jul. |
Habitat | Rocky soil | Rock outcrops and gravelly soil in meadows and open conifer forests |
Elevation | ca. 1800 m (ca. 5900 ft) | 1100-3200 m (3600-10500 ft) |
Distribution |
CA |
CA; NV; OR
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Discussion | Boechera ultra-alsa is known only from the holotype, from Snow Mountain in Lake County. Originally identified as Arabis (Boechera) platysperma, this specimen differs from that taxon by having petioles without obvious cilia, basal leaves with long-stalked (0.1–0.2 versus less than 0.1 mm) trichomes, and fruits edges parallel (not undulate or constricted between the seeds). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most authors (e.g., R. C. Rollins 1993; R. D. Dorn 2001; S. L. Welsh et al. 2003; N. H. Holmgren 2005b) have treated Boechera pinetorum as a variety of Arabis (Boechera) holboellii. Under this guise, the name has been applied to a vast array of plants collected throughout western North America. This includes a diversity of sexual diploids plus nearly every hybrid containing a genome from B. retrofracta. Based on re-examination of the type collection, we have adopted a much narrower concept of the species. Morphological evidence suggests that B. pinetorum is an apomictic triploid hybrid containing three different genomes, derived from B. rectissima, B. retrofracta, and B. sparsiflora. Plants closely resembling the type of B. pinetorum are confined to the northern Sierra Nevada and adjacent southern Cascades. The majority of collections previously associated with the epithet pinetorum represent B. pauciflora (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 410. | FNA vol. 7, p. 397. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis pinetorum, Arabis divaricarpa var. pinetorum, Arabis holboellii var. pinetorum, B. holboellii var. pinetorum | |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) | (Tidestrom) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 271. (2007) |
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