Boechera ultra-alsa |
Boechera californica |
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California rockcress |
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Habit | Perennials; long-lived; reproductive mode unknown; caudex woody. | Perennials; usually long-lived; apomictic; caudex woody. |
Stems | 1 per caudex branch, arising from center of rosette near ground surface, ca. 1 dm, densely pubescent proximally, trichomes stalked, (2 or) 3–6-rayed, 0.2–0.5 mm, glabrate distally. |
usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, (2–)3.5–13 dm, densely pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.3–0.9 mm, moderately to sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 4–6 mm wide, margins entire, not ciliate, surfaces moderately pubescent, trichomes long-stalked, 3–6 (or 7)-rayed, 0.2–0.5 mm. |
blade linear to oblanceolate, 3–10(–13) mm wide, margins entire or denticulate, ciliate along petiole, trichomes to 1.3 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.2–0.5 mm. |
Cauline leaves | 2–5, concealing stem proximally; blade auricles absent, surfaces of distalmost leaves pubescent. |
12–35(–55), often concealing stem proximally; blade auricles 1–5(–6) mm, surfaces of distalmost leaves pubescent. |
Racemes | 3- or 4-flowered, unbranched. |
30–120-flowered, usually unbranched. |
Flowers | not seen. |
divaricate-ascending at anthesis; sepals pubescent; petals usually purple, rarely pinkish, 9–14 × 1.5–3 mm, glabrous or sparsely pubescent (trichomes scattered abaxially); pollen spheroid. |
Fruiting pedicels | erect-ascending, straight, 4–5 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
descending to horizontal, usually recurved, rarely straight, 4–20 mm, pubescent, trichomes appressed, 3–7-rayed. |
Fruits | erect-ascending, sometimes appressed to rachis, not secund, straight, edges parallel, 3–4 cm × ca. 5 mm; valves glabrous; ovules ca. 16 per ovary; style 0.5–0.7 mm. |
widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (6–)8–12 cm × 1.5–2.5 mm; valves glabrous or trichomes sometimes scattered; ovules 140–180 per ovary; style 0.05–0.3 mm. |
Seeds | uniseriate, 5.5–6.5 × 4–4.5 mm; wing continuous, 1–2 mm wide. |
uniseriate or sub-biseriate, 1.7–2 × 1.2–1.7 mm; wing continuous, 0.2–0.4 mm wide. |
Boechera ultra-alsa |
Boechera californica |
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Phenology | Flowering Jun–Jul. | Flowering Mar–Jun. |
Habitat | Rocky soil | Rocky slopes and gravelly soil in desert chaparral and oak woodlands |
Elevation | ca. 1800 m (ca. 5900 ft) | 300-2300 m (1000-7500 ft) |
Distribution |
CA |
CA; Mexico (Baja California)
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Discussion | Boechera ultra-alsa is known only from the holotype, from Snow Mountain in Lake County. Originally identified as Arabis (Boechera) platysperma, this specimen differs from that taxon by having petioles without obvious cilia, basal leaves with long-stalked (0.1–0.2 versus less than 0.1 mm) trichomes, and fruits edges parallel (not undulate or constricted between the seeds). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera californica is an apomictic species that arose through hybridization between B. arcuata and B. pulchra (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). In the flora area, B. californica is known only from Los Angeles, Riverside, San Bernardino, and San Diego counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 410. | FNA vol. 7, p. 369. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis sparsiflora var. californica | |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 260. (2007) |
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