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Tulare rockcress

Pinzl's Rock cress

Habit Biennials or perennials; short-lived; apomictic; caudex present or absent. Perennials; long-lived; apomictic; caudex usually woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally.

1–3 per caudex branch, arising from margin of rosette near ground surface, 0.4–1.6 dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm.

(petiole glabrous);

blade linear-oblanceolate, 1–3 mm wide, margins entire, rarely ciliate proximally, trichomes to 0.5 mm, surfaces densely pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm.

Cauline leaves

7–17, often concealing stem proximally;

blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous.

3–6, not concealing stem;

blade auricles 0(–0.3) mm, surfaces of distalmost leaves sparsely to moderately pubescent.

Racemes

19–39-flowered, usually unbranched.

5–8-flowered, unbranched.

Flowers

divaricate to pendent at anthesis;

sepals pubescent;

petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous;

pollen spheroid.

ascending at anthesis;

sepals pubescent;

petals purple, 4–5 × 1–1.5 mm, glabrous;

pollen spheroid.

Fruiting pedicels

reflexed, recurved proximally, 5–13 mm, glabrous.

ascending to divaricate-ascending, straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

Fruits

reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm;

valves glabrous;

ovules 88–104 per ovary;

style 0.3–0.7 mm.

ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 2.5–4.8 cm × 2.5–3.2 mm;

valves glabrous;

ovules 26–34 per ovary;

style 0.1–0.3 mm.

Seeds

sub-biseriate, 2–2.5 × 1.1–1.5 mm;

wing continuous, 0.15–0.25 mm wide.

uniseriate, 2.5–3.5 × 1.5–2.5 mm;

wing continuous, 0.2–0.9 mm wide.

Boechera tularensis

Boechera pinzliae

Phenology Flowering Jun–Jul. Flowering Jul.
Habitat Rocky slopes in montane and subalpine habitats Gravelly granitic soils in alpine and subalpine areas
Elevation 2400-3200 m (7900-10500 ft) 3000-3400 m (9800-11200 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Morphological evidence suggests that Boechera pinzliae is an apomictic hybrid that contains at least one genome derived from B. platysperma; the other parent has not yet been determined. It is most similar to B. elkoensis, from which it differs in having sparsely pubescent (versus glabrous) sepals and basal leaves with stalked (versus sessile or subsessile) trichomes. Boechera pinzliae is known only from the White Mountains in extreme western Nevada (Esmeralda County) and east-central California (Mono County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 410. FNA vol. 7, p. 397.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis pinzliae
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) (Rollins) Al-Shehbaz: Novon 13: 388. (2003)
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