Boechera tularensis |
Boechera oxylobula |
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Tulare rockcress |
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Habit | Biennials or perennials; short-lived; apomictic; caudex present or absent. | Perennials; short- to long-lived; (cespitose); sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally. |
usually 3–7 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.4–2.5 dm, glabrous or pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.1–0.4 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm. |
blade linear to linear-oblanceolate, 1–2.5 mm wide, margins usually entire, rarely denticulate, often ciliate, trichomes (simple), 0.3–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2- or 3-rayed, 0.1–0.4 mm. |
Cauline leaves | 7–17, often concealing stem proximally; blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
3–12, not concealing stem; blade auricles absent, surfaces of distalmost leaves usually glabrous, rarely sparsely pubescent. |
Racemes | 19–39-flowered, usually unbranched. |
2–12-flowered, unbranched. |
Flowers | divaricate to pendent at anthesis; sepals pubescent; petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous; pollen spheroid. |
ascending-divaricate at anthesis; sepals glabrous or pubescent; petals white to pale lavender, 4–5 × 1.5–2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, recurved proximally, 5–13 mm, glabrous. |
divaricate-ascending to horizontal, slightly to strongly recurved, 3–8 mm, glabrous or sparsely pubescent, trichomes (isolated), simple. |
Fruits | reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm; valves glabrous; ovules 88–104 per ovary; style 0.3–0.7 mm. |
pendent, not appressed to rachis, not or, rarely, weakly secund, straight, edges parallel, 1.5–3.5 cm × 1.2–2 mm; valves glabrous; ovules 28–44 per ovary; style 0.1–0.4 mm. |
Seeds | sub-biseriate, 2–2.5 × 1.1–1.5 mm; wing continuous, 0.15–0.25 mm wide. |
uniseriate, 0.9–1.2 × 0.6–1 mm; wing often continuous, 0.07–0.1 mm wide. |
Boechera tularensis |
Boechera oxylobula |
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Phenology | Flowering Jun–Jul. | Flowering May–Jul. |
Habitat | Rocky slopes in montane and subalpine habitats | Cliffs, rocky slopes, gravelly soil in sagebrush and open conifer forests |
Elevation | 2400-3200 m (7900-10500 ft) | 2100-3600 m (6900-11800 ft) |
Distribution |
CA |
CO |
Discussion | Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most of the collections assigned here have been called Arabis (Boechera) demissa by other authors (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b). Because the holotype of A. demissa is identical to B. oxylobula in nearly every way (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison), we treat them as conspecific. Boechera oxylobula is restricted to Garfield, Gunnison, Hinsdale, Lake, Mineral, Park, and Saguache counties in central Colorado. The taxon traditionally treated as A. (Boechera) demissa var. languida is here recognized as an apomictic species of hybrid origin (see Windham and Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 410. | FNA vol. 7, p. 391. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis oxylobula, Arabis aprica, Arabis demissa, Arabis rugocarpa, B. demissa | |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) | (Greene) W. A. Weber: Phytologia 51: 370. (1982) |
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