Boechera tularensis |
Boechera johnstonii |
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Tulare rockcress |
Johnston's rockcress |
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Habit | Biennials or perennials; short-lived; apomictic; caudex present or absent. | Perennials; long-lived; (cespitose); sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2 dm, densely pubescent proximally, trichomes short-stalked, 4–10-rayed, 0.07–0.15 mm, sparsely to densely pubescent distally. |
Basal leaves | blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm. |
blade narrowly oblanceolate, 1.5–4 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 6–14-rayed, 0.07–0.15 mm. |
Cauline leaves | 7–17, often concealing stem proximally; blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
4–10, often concealing stem proximally; blade auricles absent, surfaces of distalmost leaves densely pubescent. |
Racemes | 19–39-flowered, usually unbranched. |
10–18-flowered, unbranched. |
Flowers | divaricate to pendent at anthesis; sepals pubescent; petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, recurved proximally, 5–13 mm, glabrous. |
divaricate-ascending, straight, 5–14 mm, pubescent, trichomes appressed, branched. |
Fruits | reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm; valves glabrous; ovules 88–104 per ovary; style 0.3–0.7 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 4–6 cm × 2.5–4 mm; valves glabrous; ovules 26–34 per ovary; style (0.7–)1–2 mm. |
Seeds | sub-biseriate, 2–2.5 × 1.1–1.5 mm; wing continuous, 0.15–0.25 mm wide. |
uniseriate, 1.9–2.7 × 1.5–2.2 mm; wing continuous, 0.3–0.7 mm wide. |
Boechera tularensis |
Boechera johnstonii |
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Phenology | Flowering Jun–Jul. | Flowering Feb–Mar. |
Habitat | Rocky slopes in montane and subalpine habitats | Rocky areas and gravelly soil in chaparral and oak-pine savannas |
Elevation | 2400-3200 m (7900-10500 ft) | 1300-1700 m (4300-5600 ft) |
Distribution |
CA |
CA |
Discussion | Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Boechera johnstonii and B. hirshbergiae are too similar to be considered distinct species. Even with the expanded circumscription, this distinctive sexual species is known only from the area of Cuyamaca Lake (San Diego County) and the San Jacinto Mountains (Riverside County). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 410. | FNA vol. 7, p. 386. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis johnstonii, Arabis hirshbergiae, B. hirshbergiae | |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) | (Munz) Al-Shehbaz: Novon 13: 386. (2003) |
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