Boechera subpinnatifida |
Boechera xylopoda |
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ashy rock-cress, Klamath rockcress |
beautiful rockcress, bigfoot hybrid rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 from per caudex branch, arising from center of leaf tuft, elevated above ground surface on woody base, (3–)5–7.5 dm, pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.25–0.9 mm., similarly pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade linear-oblanceolate to oblanceolate, 3–7 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 1.5 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.15–0.5 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
8–18, usually not concealing stem; blade auricles 1–2 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
20–38-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals purple, 9–12 × 2–3.5 mm, sparsely pubescent (often some trichomes abaxially); pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
descending-divaricate, usually recurved, 9–20 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
reflexed, not appressed to rachis, not secund, straight, edges parallel, 5–7 cm × 2–2.5 mm; valves glabrous proximally, sparsely pubescent distally; ovules 98–126 per ovary; style 0.1–0.5 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
sub-biseriate, 2–2.5 × 1.3–1.6 mm; wing continuous, 0.2–0.25 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera xylopoda |
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Phenology | Flowering Mar–May. | Flowering Mar–May. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rock outcrops and gravelly slopes, often under shelter of shrubs in desert scrub, sagebrush, and chaparral communities |
Elevation | 800-2400 m (2600-7900 ft) | 900-2000 m (3000-6600 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; NV
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although usually treated as a variety of Arabis (Boechera) pulchra, B. xylopoda is an apomictic taxon that is morphologically intermediate between B. perennans and B. pulchra (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison) and probably arose through hybridization between those two sexual diploids. Arabis trichopoda Greene (1908), not Turczaninow (1840) is an illegitimate name, sometimes found in synonymy with Boechera xylopoda. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 411. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis pulchra var. gracilis, Arabis pulchra var. glabrescens, Arabis pulchra var. viridis, B. pulchra var. gracilis |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 250. (2007) |
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