Boechera subpinnatifida |
Boechera villosa |
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ashy rock-cress, Klamath rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
1–3 per caudex branch, arising laterally proximal to sterile shoots near ground surface, ca. 2.5 dm, densely pubescent proximally, trichomes simple and 2-rayed, 0.25–0.7 mm, glabrescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate to obovate, 2.5–5 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes simple and short-stalked, 2- or 3-rayed, 0.25–0.5 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
4–6, not concealing stem; blade auricles ca. 1 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
6–10-flowered, unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender, 4–5 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
ascending, straight, 6–10 mm, glabrous or sparsely pubescent, trichomes subappressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
divaricate-ascending, not appressed to rachis, not secund, slightly curved, edges parallel, 4–5 cm × ca. 1 mm; valves glabrous; ovules ca. 64 per ovary; style ca. 0.2 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, (none mature). |
2n | = 14. |
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Boechera subpinnatifida |
Boechera villosa |
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Phenology | Flowering Mar–May. | Flowering May. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Basalt outcrop in pinyon-juniper woodlands |
Elevation | 800-2400 m (2600-7900 ft) | ca. 2100 m (ca. 6900 ft) |
Distribution |
CA; ID; NV; OR; UT
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NM |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera villosa is known only from the type collection from the Rio Grande Gorge in Taos County. The holotype was originally identified as Arabis (Boechera) perennans but clearly is more closely related to B. pallidifolia (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 411. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) |
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