Boechera subpinnatifida |
Boechera pratincola |
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ashy rock-cress, Klamath rockcress |
meadow rock-cress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; usually short-lived; apomictic; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6 dm, pubescent proximally, trichomes sessile, 2- or 3-rayed, 0.15–0.6 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate, 1.5–5(–7) mm wide, margins usually entire, rarely denticulate, ciliate at petiole base, trichomes to 0.4 mm, surfaces pubescent, trichomes sessile, 2–4(–6)-rayed, 0.1–0.45 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
(7–)13–42, concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves sparsely pubescent apically. |
Racemes | 8–30-flowered, usually unbranched. |
(7–)14–45-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous; petals lavender, 6–10 × 1.5–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
erect, straight, 5–12 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
erect, appressed to rachis, not secund, straight, edges parallel, 4–6.5 cm × 1.5–2 mm; valves glabrous; ovules 60–140 per ovary; style 0.1–0.3 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
sub-biseriate, 1.4–2 × 1–1.4 mm; wing continuous, 0.1–0.35 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera pratincola |
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Phenology | Flowering Mar–May. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and soil patches in open areas and along forest edges |
Elevation | 800-2400 m (2600-7900 ft) | 1900-3200 m (6200-10500 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; ID; NV; OR
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera pratincola is an apomictic species that arose through hybridization between B. paupercula and B. stricta (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 399. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis pratincola, Arabis drummondii var. pratincola, Arabis nemophila |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 245. (2007) |
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