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Boechera subpinnatifida

ashy rock-cress, Klamath rockcress

Pinzl's Rock cress

Habit Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). Perennials; long-lived; apomictic; caudex usually woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally.

1–3 per caudex branch, arising from margin of rosette near ground surface, 0.4–1.6 dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm.

(petiole glabrous);

blade linear-oblanceolate, 1–3 mm wide, margins entire, rarely ciliate proximally, trichomes to 0.5 mm, surfaces densely pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm.

Cauline leaves

(10–)20–60, often concealing stem throughout;

blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent.

3–6, not concealing stem;

blade auricles 0(–0.3) mm, surfaces of distalmost leaves sparsely to moderately pubescent.

Racemes

8–30-flowered, usually unbranched.

5–8-flowered, unbranched.

Flowers

divaricate-ascending to pendent at anthesis;

sepals pubescent;

petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals purple, 4–5 × 1–1.5 mm, glabrous;

pollen spheroid.

Fruiting pedicels

reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched.

ascending to divaricate-ascending, straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

Fruits

pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm;

valves pubescent throughout;

ovules 24–42 per ovary;

style 0.5–1 mm.

ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 2.5–4.8 cm × 2.5–3.2 mm;

valves glabrous;

ovules 26–34 per ovary;

style 0.1–0.3 mm.

Seeds

uniseriate, 2.5–3.5 × 1.5–2.2 mm;

wing continuous or at both ends, 0.4–0.8 mm wide.

uniseriate, 2.5–3.5 × 1.5–2.5 mm;

wing continuous, 0.2–0.9 mm wide.

2n

= 14.

Boechera subpinnatifida

Boechera pinzliae

Phenology Flowering Mar–May. Flowering Jul.
Habitat Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities Gravelly granitic soils in alpine and subalpine areas
Elevation 800-2400 m (2600-7900 ft) 3000-3400 m (9800-11200 ft)
Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Morphological evidence suggests that Boechera pinzliae is an apomictic hybrid that contains at least one genome derived from B. platysperma; the other parent has not yet been determined. It is most similar to B. elkoensis, from which it differs in having sparsely pubescent (versus glabrous) sepals and basal leaves with stalked (versus sessile or subsessile) trichomes. Boechera pinzliae is known only from the White Mountains in extreme western Nevada (Esmeralda County) and east-central California (Mono County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 409. FNA vol. 7, p. 397.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis subpinnatifida Arabis pinzliae
Name authority (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) (Rollins) Al-Shehbaz: Novon 13: 388. (2003)
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