Boechera subpinnatifida |
Boechera lincolnensis |
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ashy rock-cress, Klamath rockcress |
Lincoln rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of leaf tuft, elevated above ground surface on woody base, 2–4.2 dm, densely pubescent proximally, trichomes short-stalked, 3–6-rayed, 0.1–0.5 mm, similarly pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade linear or linear-oblanceolate, 1–2 mm wide, margins entire, rarely ciliate near petiole base, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 3–8-rayed, 0.1–0.4 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
10–25, concealing stem proximally; blade auricles absent or, rarely, to 1 mm, surfaces of distalmost leaves densely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
7–15-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender to purple, 10–12 × 2–3 mm, glabrous or sparsely pubescent (occasionally some trichomes abaxially); pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending, straight to slightly curved proximally, recurved distally, 10–20(–25) mm, pubescent, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
usually widely pendent to ascending, rarely horizontal, not appressed to rachis, not secund, straight, edges parallel, (3.2–)4–5.5 cm × 2–2.5 mm; valves pubescent throughout; ovules 86–120 per ovary; style 0.1–0.3 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
biseriate, 1–1.5 × 0.7–0.9 mm; wing continuous, 0.07–0.12 mm wide. |
2n | = 14. |
= 14. |
Boechera subpinnatifida |
Boechera lincolnensis |
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Phenology | Flowering Mar–May. | Flowering Apr–May. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and gravelly soil with sagebrush and other shrubs |
Elevation | 800-2400 m (2600-7900 ft) | 1400-1900 m (4600-6200 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; NV; UT |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Recent studies suggest that Boechera lincolnensis is a distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 388. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis pulchra var. munciensis, B. pulchra var. munciensis |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 71. (2006) |
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