Boechera subpinnatifida |
Boechera grahamii |
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ashy rock-cress, Klamath rockcress |
Graham's rockcress, Graham's suncress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.2–)2–12 dm, glabrous or pubescent proximally, trichomes simple and sessile or subsessile, 2 or 3(–6)-rayed, to 1 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate, 1.5–10(–20) mm wide, margins entire or dentate, ciliate proximally, trichomes (simple), to 1 mm, surfaces sparsely to densely pubescent, trichomes sessile or subsessile, 2–4(–7)-rayed, 0.2–0.6 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
(10–)13–52, often concealing stem proximally; blade auricles 1–5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
16–88-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
divaricate at anthesis; sepals glabrous or pubescent; petals usually white, rarely lavender, 5.5–8 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending to descending, usually gently curved downward, rarely straight, 5–22(–30) mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
divaricate-ascending to pendent, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 3.5–9 cm × 1–1.8 mm; valves glabrous; ovules 84–146 per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate to sub-biseriate, 1–1.8 × 0.8–1.2 mm; wing continuous, 0.1–0.25 mm wide. |
2n | = 14. |
= 21. |
Boechera subpinnatifida |
Boechera grahamii |
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Phenology | Flowering Mar–May. | Flowering May–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and sandy soil in prairies and open forests |
Elevation | 800-2400 m (2600-7900 ft) | 100-3200 m (300-10500 ft) |
Distribution |
CA; ID; NV; OR; UT
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AK; CO; MI; MN; MT; ND; NY; SD; VT; WA; WI; WY; AB; BC; MB; NT; ON; QC; SK; YT
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera grahamii is an apomictic species that arose through hybridization between B. collinsii and B. stricta. Previous authors have assigned these specimens to Arabis (Boechera) divaricarpa (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison); if these taxa are treated as conspecific, the name B. grahamii has priority. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 382. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Turritis grahamii, Arabis boivinii, Arabis bourgovii, Arabis confinis, Arabis confinis var. brachycarpa, Arabis dacotica, Arabis divaricarpa var. dacotica, Arabis divaricarpa var. hemicylindrica, Arabis divaricarpa var. stenocarpa, Arabis drummondii var. brachycarpa, Arabis holboellii var. brachycarpa, B. brachycarpa, Turritis brachycarpa |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Lehmann) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 241. (2007) |
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