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Boechera subpinnatifida

ashy rock-cress, Klamath rockcress

Habit Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). Perennials; short- to long-lived; sexual; caudex usually not woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally.

usually 2–6 per caudex branch, arising from margin of rosette near ground surface, 0.8–4 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.3–0.6 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm.

blade narrowly oblanceolate, 1–5 mm wide, margins entire or few-toothed, ciliate proximally, trichomes (simple), to 0.9 mm, surfaces densely pubescent, trichomes simple and short-stalked, 2- or 3-rayed, 0.3–0.5 mm.

Cauline leaves

(10–)20–60, often concealing stem throughout;

blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent.

3–6, not concealing stem;

blade auricles 0.3–0.6 mm, surfaces of distalmost leaves glabrous.

Racemes

8–30-flowered, usually unbranched.

7–15-flowered, usually unbranched.

Flowers

divaricate-ascending to pendent at anthesis;

sepals pubescent;

petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals lavender, 3.5–6 × 1–1.5 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched.

divaricate-ascending to horizontal, straight or gently recurved, 4–10 mm, glabrous or sparsely pubescent, trichomes spreading.

Fruits

pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm;

valves pubescent throughout;

ovules 24–42 per ovary;

style 0.5–1 mm.

divaricate-ascending to horizontal, not appressed to rachis, not secund, usually curved or straight, edges parallel, 2.5–4 cm × 1.2–1.5 mm;

valves glabrous;

ovules 50–80 per ovary;

style 0.05–0.3 mm.

Seeds

uniseriate, 2.5–3.5 × 1.5–2.2 mm;

wing continuous or at both ends, 0.4–0.8 mm wide.

uniseriate, 1–1.5 × 0.7–1 mm;

wing usually distal, ca. 0.1 mm wide.

2n

= 14.

Boechera subpinnatifida

Boechera glareosa

Phenology Flowering Mar–May. Flowering May–Jun.
Habitat Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities Conglomerate and limestone outcrops in pinyon-juniper and mountain shrub-conifer communities
Elevation 800-2400 m (2600-7900 ft) 2000-2600 m (6600-8500 ft)
Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; UT
[BONAP county map]
Discussion

Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Specimens of Boechera glareosa originally were identified as Arabis microphylla var. macounii (= B. macounii) by R. C. Rollins. The former (a sexual species) is distinguished from the latter (an apomictic hybrid) by having wider (1–2 versus 0.5–0.8 mm) petals, fewer (2–9 versus mostly 9–16) cauline leaves, and fewer (7–15 versus 10–33) flowers per inflorescence. Boechera glareosa is known only from Moffat County, Colorado, and Summit and Uintah counties, Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 409. FNA vol. 7, p. 380.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis subpinnatifida
Name authority (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) Dorn: Brittonia 55: 1, fig. 1. (2003)
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