Boechera subpinnatifida |
Boechera evadens |
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ashy rock-cress, Klamath rockcress |
hidden rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; (cespitose); sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1–2.5 dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.2–0.6 mm, and sparse, simple ones to 1 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade narrowly oblanceolate, 1.5–3 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.2 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
3–5, not concealing stem; blade auricles 0.5–1 mm, surfaces of distalmost leaves pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
10–22-flowered, unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending, straight, 4–8 mm, sparsely pubescent, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 3–4 cm × 1.2–1.5 mm; valves glabrous; ovules 46–52 per ovary; style ca. 0.1 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1–1.1 × 0.9–1 mm; wing continuous, ca. 0.1 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera evadens |
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Phenology | Flowering Mar–May. | Flowering Jun. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rock outcrops |
Elevation | 800-2400 m (2600-7900 ft) | ca. 2600 m (ca. 8500 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera evadens is a sexual taxon that was originally interpreted as a disjunct population of Arabis (Boechera) fernaldiana, but is easily distinguished from that species by having much shorter (3–4 versus 7–12 mm) petals, larger (0.2–0.6 versus 0.04–0.1 mm) trichomes proximally on stems, and shorter (3–4 versus 4–7.5 cm) fruits (see M. D. Windham and I. A. Al-Shehbaz 2006 for additional distinctions). It is known only from the holotype specimen collected near Sherman Pass in Tulare County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 376. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 66. (2006) |
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