Boechera subpinnatifida |
Boechera drepanoloba |
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ashy rock-cress, Klamath rockcress |
pointing suncress, soldier rock-cress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; short- to long-lived; apomictic; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4 dm, glabrous or sparsely pubescent proximally, trichomes sessile or subsessile, 2- or 3-rayed, 0.1–0.3 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate, 2–6 mm wide, margins entire, ciliate along petiole, trichomes (submalpighiaceous or simple), to 0.6 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–6-rayed, 0.1–0.3 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
3–15, often concealing stem proximally; blade auricles 0.7–2.5 mm, surfaces of distalmost leaves usually glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
5–25-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous or pubescent with scattered trichomes; petals purple to lavender, 6–8 × 1.5–2.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
horizontal to ascending, straight or curved, 4–8 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
horizontal to ascending, secund, straight to slightly curved, edges parallel, 3–5.8 cm × 2–3.5 mm; valves glabrous; ovules 44–76(–104) per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1.5–2.2 × 1.2–1.6 mm; wing continuous, 0.3–0.7 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera drepanoloba |
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Phenology | Flowering Mar–May. | Flowering Jul–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and talus in alpine and subalpine habitats |
Elevation | 800-2400 m (2600-7900 ft) | 2000-3800 m (6600-12500 ft) |
Distribution |
CA; ID; NV; OR; UT
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AK; CO; ID; MT; OR; UT; WA; WY; AB |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera drepanoloba is an apomictic species that arose through hybridization between B. lemmonii and B. stricta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 374. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis drepanoloba, Arabis drummondii var. oreophila, Arabis lemmonii var. drepanoloba, Arabis oreophila |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 263. (2007) |
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