Boechera subpinnatifida |
Boechera depauperata |
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ashy rock-cress, Klamath rockcress |
soldier rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; usually long-lived; (cespitose); apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–1.5(–2) dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.07–0.4 mm, usually sparsely pubescent distally, rarely glabrous. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade narrowly oblanceolate, 0.7–3(–5) mm wide, margins entire, sometimes ciliate proximally, trichomes to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 5–8-rayed, 0.05–0.25 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves usually pubescent, rarely glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
9–23-flowered, unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender to purplish, 3.5–5 × 0.8–1.2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending, straight, 1.5–7 mm, usually pubescent, rarely glabrous, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 1.7–3.5(–4.5) cm × 1–1.5 mm; valves glabrous; ovules 32–44 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1.2–1.5 × 1–1.2 mm; wing continuous, 0.1–0.15 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera depauperata |
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Phenology | Flowering Mar–May. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Exposed ridges and talus slopes in subalpine and alpine habitats |
Elevation | 800-2400 m (2600-7900 ft) | 2700-3900 m (8900-12800 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; NV
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera depauperata is an apomictic species that arose through hybridization between B. lemmonii and B. paupercula (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). It is known in California from Fresno, Mono, and Tulare counties and in Nevada from Washoe County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 373. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis depauperata, Arabis lemmonii var. depauperata |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (A. Nelson & P. B. Kennedy) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 262. (2007) |
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