Boechera subpinnatifida |
Boechera consanguinea |
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ashy rock-cress, Klamath rockcress |
Four Corners rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–5 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.2–0.5 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate, 2–10 mm wide, margins usually dentate, sometimes ciliate near petiole base, trichomes (usually spurred), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.4 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
15–36, sometimes concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
20–55-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
divaricate-descending at anthesis; sepals pubescent; petals pale lavender, 5–8.5 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
reflexed to divaricate-descending, usually curved downward, 8–14 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
reflexed to pendent, rarely appressed to rachis, not secund, straight or slightly curved, edges parallel, 4–6 cm × 1–2 mm; valves glabrous; ovules 100–128 per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
biseriate to sub-biseriate, mature seeds not seen. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera consanguinea |
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Phenology | Flowering Mar–May. | Flowering May. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and sandy soil in ponderosa pine, pinyon-juniper, and sagebrush communities |
Elevation | 800-2400 m (2600-7900 ft) | 1900-2500 m (6200-8200 ft) |
Distribution |
CA; ID; NV; OR; UT
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CO; NM; UT
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera consanguinea is an apomictic species that arose through hybridization between B. fendleri and B. retrofracta. It is most similar to B. goodrichii, another apomictic hybrid involving B. retrofracta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 371. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis consanguinea, Arabis holboellii var. consanguinea |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 261. (2007) |
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