Boechera subpinnatifida |
Boechera calderi |
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ashy rock-cress, Klamath rockcress |
Calder's rock-cress, Calder's suncress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; usually short-lived; apomictic; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4.5 dm, glabrous throughout. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate, 1.5–6 mm wide, margins entire, ciliate at petiole base, trichomes to 0.8 mm, surfaces sparsely pubescent, trichomes sessile, malpighiaceous (usually at margin) and 3- or 4-rayed, 0.15–0.4 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
5–17, often concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
10–25-flowered, unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
erect at anthesis; sepals glabrous; petals purple, 6–9 × 1.5–2.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
erect, straight, 5–10 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
erect, appressed to rachis, not secund, straight, edges parallel, 3.5–6.5 cm × 1.8–2.5 mm; valves glabrous; ovules 64–134 per ovary; style 0–0.2 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
biseriate to sub-biseriate, 1.8–2.2 × 1–1.8 mm; wing continuous, 0.2–0.5 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera calderi |
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Phenology | Flowering Mar–May. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Exposed rocky ridges, meadows, open forests near timberline |
Elevation | 800-2400 m (2600-7900 ft) | 1500-3500 m (4900-11500 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; ID; MT; OR; WA; WY; AB; BC; NT; YT |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera calderi is an apomictic species that arose through hybridization between B. lyallii and B. stricta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 368. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis calderi |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (G. A. Mulligan) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 259. (2007) |
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