Boechera subpinnatifida |
Boechera burkii |
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ashy rock-cress, Klamath rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Biennials; short-lived; sexual; caudex not evident. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per plant, arising from center of rosette near ground surface, (2–)3–7 dm, glabrous throughout, often glaucous. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate to obovate, 4–10 mm wide, margins dentate, not ciliate, surfaces glabrous or sparsely pubescent subapically, trichomes simple, 0.3–0.8 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
18–28, often concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
20–50-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous; petals white, 3–5 × 0.5–0.7 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
ascending to divaricate-ascending, usually straight, 4–12 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 5–10 cm × 1.5–1.8 mm; valves glabrous; ovules 64–80 per ovary; style 0.2–0.6 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1.5–1.8 × 1.2–1.4 mm; wing continuous, 0.3–0.5 mm wide distally. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera burkii |
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Phenology | Flowering Mar–May. | Flowering Apr–May. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky areas, wooded slopes, stream banks |
Elevation | 800-2400 m (2600-7900 ft) | |
Distribution |
CA; ID; NV; OR; UT
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MD; PA; TN; VA; WV |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera burkii usually has been treated as a variety of Arabis laevigata (e.g., R. C. 1993). The two taxa differ substantially in leaf morphology, and no intermediates have been found despite broad overlap in their geographic ranges. Boechera burkii typically has 18–28 cauline leaves per plant, with blades linear, non-auriculate, and margins entire, whereas B. laevigata has 7–15 cauline leaves per plant, blades lanceolate, auriculate, and margins often dentate. The two are also separable on petal width, with B. burkii having significantly narrower petals (0.5–0.7 versus 1–1.5 mm wide). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 368. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis laevigata var. burkii, Arabis burkii |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Porter) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 237. (2007) |
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