Boechera subpinnatifida |
Boechera bodiensis |
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ashy rock-cress, Klamath rockcress |
Bodie Hills rock cress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; (cespitose); apomictic; caudex somewhat woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–3.5 dm, densely pubescent proximally, trichomes short-stalked, 5–10-rayed, to 0.7 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate proximally, trichomes to 0.7 mm, surfaces densely pubescent, trichomes short-stalked, 5–10-rayed, 0.08–0.25 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
4–9, not concealing stem; blade auricles 0.5–2 mm, surfaces of distalmost leaves pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
8–25-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender to purple, 4–5 × ca. 1 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
ascending to divaricate-ascending, straight, 3–7 mm, pubescent, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 4–6.2 cm × 1.2–1.8 mm; valves glabrous; ovules 48–68 per ovary; style 0.1–0.2 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1–1.5 × 0.8–1.2 mm; wing continuous, 0.1–0.15 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera bodiensis |
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Phenology | Flowering Mar–May. | Flowering Jul–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Loose soil, crevices of igneous rock |
Elevation | 800-2400 m (2600-7900 ft) | 2400-2900 m (7900-9500 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; NV |
Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. In his original description, Rollins suggested that Boechera bodiensis is closely related to B. cobrensis and B. falcifructa. Subsequent studies (M. D. Windham and I. A. Al-Shehbaz, unpubl.) indicate that both B. bodiensis and B. falcifructa are apomictic hybrids containing one or more genomes derived from B. cobrensis. In the case of B. falcifructa, the second parent is clearly B. fernaldiana, but the second parent of B. bodiensis remains uncertain. Boechera bodiensis is most similar to B. falcifructa. The latter differs by having gently recurved fruiting pedicels 6–12 mm, 62–80 ovules per ovary, and pendulous fruits; it is known from Elko and Lander counties, Nevada. By contrast, B. bodiensis has straight fruiting pedicels 3–7 mm long, divaricate-ascending fruits, 48–68 ovules per ovary, and is known from Mono County, California, and Lyon and Mineral counties, Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 367. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis bodiensis |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Rollins) Al-Shehbaz: Novon 13: 384. (2003) |
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