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Boechera subpinnatifida

ashy rock-cress, Klamath rockcress

point rockcress, sharp-pod rock-cress

Habit Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). Biennials or perennials; short-lived; apomictic; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–6 dm, densely pubescent proximally, trichomes sessile, 2–4-rayed, 0.15–0.4 mm, usually sparsely pubescent distally, rarely glabrous.

Basal leaves

blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm.

blade oblanceolate, 1.5–6 mm wide, margins usually entire, rarely denticulate, ciliate proximally, trichomes (simple), to 0.5 mm, surfaces sparsely to moderately pubescent, trichomes sessile, 3–5 (or 6)-rayed, 0.15–0.35 mm.

Cauline leaves

(10–)20–60, often concealing stem throughout;

blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent.

2–20(–38), often concealing stem proximally;

blade auricles 0.3–2 mm, surfaces of distalmost leaves sparsely pubescent or glabrous.

Racemes

8–30-flowered, usually unbranched.

5–20(–36)-flowered, usually unbranched.

Flowers

divaricate-ascending to pendent at anthesis;

sepals pubescent;

petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals white or lavender, 5–7 × 0.9–1.3 mm, glabrous;

pollen spheroid.

Fruiting pedicels

reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched.

erect to ascending, usually straight, 3–10 mm, pubescent or glabrous, trichomes appressed, branched.

Fruits

pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm;

valves pubescent throughout;

ovules 24–42 per ovary;

style 0.5–1 mm.

ascending, not appressed to rachis, not secund, straight, edges parallel, 2.5–7.5 cm × 1.2–1.8 mm;

valves glabrous;

ovules 46–100 per ovary;

style 0.3–1 mm.

Seeds

uniseriate, 2.5–3.5 × 1.5–2.2 mm;

wing continuous or at both ends, 0.4–0.8 mm wide.

uniseriate, 1.8–2.2 × 1–1.4 mm;

wing continuous, 0.1–0.4 mm wide.

2n

= 14.

Boechera subpinnatifida

Boechera acutina

Phenology Flowering Mar–May. Flowering Jun–Jul.
Habitat Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities Gravelly slopes in meadows, open forests
Elevation 800-2400 m (2600-7900 ft) 1400-1900 m (4600-6200 ft)
Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; OR
[BONAP county map]
Discussion

Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera acutina is an apomictic hybrid that clearly contains a genome derived from B. stricta; the other parent remains uncertain. Plants of B. acutina are superficially similar to B. pratincola and are often misidentified as Arabis (Boechera) divaricarpa; the species is easily distinguished from both of those (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 409. FNA vol. 7, p. 365.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis subpinnatifida Arabis acutina, Arabis divaricarpa var. interposita, Arabis drummondii var. interposita, Arabis interposita
Name authority (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 236. (2007)
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