Boechera pygmaea |
Boechera spatifolia |
|
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Tulare County rock cress |
spoonleaf rockcress |
|
Habit | Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases). | Perennials; short- to long-lived; sexual; caudex usually not woody (rarely with persistent, crowded leaf bases). |
Stems | usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally. |
simple or few to several per caudex branch, arising from center of rosette near ground surface, 1.5–3.5(–5) dm, densely pubescent proximally, trichomes simple mixed with short-stalked ones, 2-rayed, 0.3–0.7 mm. |
Basal leaves | blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm. |
blade narrowly oblanceolate, 1.5–3(–4) mm wide, margins entire, strongly ciliate at least along petiole, trichomes (simple), to 1 mm, surfaces glabrous or sparsely pubescent, trichomes simple and short-stalked, 2-rayed, 0.3–0.7 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles absent, surfaces of distalmost leaves pubescent. |
5–15(–20), often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves usually glabrous. |
Racemes | 2–5-flowered, unbranched. |
10–30-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals pubescent; petals white, 3.5–5 × 0.7–1 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals usually white, rarely pale lavender, 3–3.7(–4) × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
horizontal or slightly descending, curved or angled downward, 6–10(–15) mm, usually glabrous, rarely sparsely pubescent, trichomes spreading, simple. |
Fruits | erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm; valves glabrous; ovules 8–12 per ovary; style 0.05–0.4 mm. |
pendent, not appressed to rachis, rarely slightly secund, straight or gently curved, edges parallel, 3.3–5.7 cm × 1.2–1.8 mm; valves glabrous; ovules 90–126 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate, 3–5 × 2.5–4.5 mm; wing continuous, 0.8–2 mm wide distally. |
biseriate, 0.7–0.9 × 0.5–0.6 mm; wing distal or, sometimes, absent, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera pygmaea |
Boechera spatifolia |
|
Phenology | Flowering May–Jul. | Flowering May–Aug. |
Habitat | Barren flats of arkosic gravel | Rocky slopes and gravelly soil in sagebrush, pinyon-juniper woodlands, open conifer forests and subalpine meadows |
Elevation | 2400-3200 m (7900-10500 ft) | 1800-2800 m (5900-9200 ft) |
Distribution |
CA |
CO; NM
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Discussion | Of conservation concern. Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera spatifolia is a sexual diploid that usually has been treated as a variety of Arabis (Boechera) fendleri but appears to be sufficiently distinct to warrant recognition at species level (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). There is little geographic overlap between the two, with B. spatifolia confined to the mountains of central Colorado and north-central New Mexico and B. fendleri ranging from western New Mexico and the Four Corners region through northern Arizona to southern Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 401. | FNA vol. 7, p. 408. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis pygmaea | Arabis spatifolia, Arabis fendleri var. spatifolia, B. fendleri subsp. spatifolia, B. fendleri var. spatifolia |
Name authority | (Rollins) Al-Shehbaz: Novon 13: 388. (2003) | (Rydberg) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 84. (2006) |
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