Boechera pygmaea |
Boechera shockleyi |
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Tulare County rock cress |
Shockley's Rock cress |
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Habit | Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases). | Perennials; usually short-lived; sexual; caudex not woody, (rarely with persistent, crowded leaf bases). |
Stems | usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (0.8–)2–5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
Basal leaves | blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm. |
blade oblanceolate, 3–10 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles absent, surfaces of distalmost leaves pubescent. |
14–60, concealing stem for most of length; blade auricles 0.5–4 mm, surfaces of distalmost leaves densely pubescent. |
Racemes | 2–5-flowered, unbranched. |
20–70-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals pubescent; petals white, 3.5–5 × 0.7–1 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender, 6–9 × 0.8–1.2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
divaricate-ascending, straight, 7–28 mm, pubescent, trichomes appressed, branched. |
Fruits | erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm; valves glabrous; ovules 8–12 per ovary; style 0.05–0.4 mm. |
divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 4.5–11 cm × 1.5–2 mm; valves glabrous or sparsely pubescent throughout; ovules 140–190 per ovary; style 0.05–0.6 mm. |
Seeds | uniseriate, 3–5 × 2.5–4.5 mm; wing continuous, 0.8–2 mm wide distally. |
sub-biseriate, 1–1.3 × 0.7–0.8 mm; wing distal or, rarely, absent, 0.05–0.1 mm wide. |
Boechera pygmaea |
Boechera shockleyi |
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Phenology | Flowering May–Jul. | Flowering Apr–May. |
Habitat | Barren flats of arkosic gravel | Rock outcrops (primarily dolomite) and gravelly soil in desert scrub, sagebrush, and pinyon-juniper woodlands |
Elevation | 2400-3200 m (7900-10500 ft) | 1200-2200 m (3900-7200 ft) |
Distribution |
CA |
CA; NV; UT
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Discussion | Of conservation concern. Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera shockleyi is a distinctive sexual species recognizable by the combination of strongly overlapping cauline leaves, a dense covering of minute, 7–12-rayed trichomes, and relatively long, curved, divaricate-ascending fruits with sub-biseriate seeds. It is most similar to B. inyoensis, but differs from that species in its narrower (0.8–1.2 versus 1.2–2 mm) petals, eciliate basal leaves with smaller (0.1–0.2 versus 0.2–0.7 mm) trichomes, greater number (140–190 versus 74–134) of ovules per ovary, and shorter (1–1.3 versus 1.7–2 mm) seeds with distal (versus continuous) wings. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 401. | FNA vol. 7, p. 406. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis pygmaea | Arabis shockleyi |
Name authority | (Rollins) Al-Shehbaz: Novon 13: 388. (2003) | (Munz) Dorn: Brittonia 55: 3. (2003) |
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