Boechera pygmaea |
Boechera pinetorum |
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Tulare County rock cress |
Holboell's rock-cress, woodland rock-cress |
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Habit | Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases). | Perennials; short-lived; apomictic; caudex not woody. |
Stems | usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–8(–9.6) dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm. |
blade oblanceolate, 2–8(–11) mm wide, margins entire or denticulate, ciliate proximally, trichomes (simple), 0.5–1 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed, 0.2–0.5 mm, these mixed with fewer simple ones. |
Cauline leaves | 2–4, not concealing stem; blade auricles absent, surfaces of distalmost leaves pubescent. |
7–33, often concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
Racemes | 2–5-flowered, unbranched. |
15–63-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals pubescent; petals white, 3.5–5 × 0.7–1 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals white to lavender, 5–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
reflexed, abruptly recurved near base, otherwise straight, 4–12 mm, sparsely pubescent or glabrous, trichomes spreading, simple and 2-rayed. |
Fruits | erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm; valves glabrous; ovules 8–12 per ovary; style 0.05–0.4 mm. |
reflexed to closely pendent, not appressed to rachis, not secund, straight or curved, edges parallel, (4.5–)5.5–8.5 cm × 1.5–2 mm; valves glabrous; ovules 70–110 per ovary; style 0.05–0.4 mm. |
Seeds | uniseriate, 3–5 × 2.5–4.5 mm; wing continuous, 0.8–2 mm wide distally. |
uniseriate, 1.5–1.8 × 1.2–1.5 mm; wing continuous, 0.15–0.3 mm wide. |
Boechera pygmaea |
Boechera pinetorum |
|
Phenology | Flowering May–Jul. | Flowering May–Jul. |
Habitat | Barren flats of arkosic gravel | Rock outcrops and gravelly soil in meadows and open conifer forests |
Elevation | 2400-3200 m (7900-10500 ft) | 1100-3200 m (3600-10500 ft) |
Distribution |
CA |
CA; NV; OR
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Discussion | Of conservation concern. Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most authors (e.g., R. C. Rollins 1993; R. D. Dorn 2001; S. L. Welsh et al. 2003; N. H. Holmgren 2005b) have treated Boechera pinetorum as a variety of Arabis (Boechera) holboellii. Under this guise, the name has been applied to a vast array of plants collected throughout western North America. This includes a diversity of sexual diploids plus nearly every hybrid containing a genome from B. retrofracta. Based on re-examination of the type collection, we have adopted a much narrower concept of the species. Morphological evidence suggests that B. pinetorum is an apomictic triploid hybrid containing three different genomes, derived from B. rectissima, B. retrofracta, and B. sparsiflora. Plants closely resembling the type of B. pinetorum are confined to the northern Sierra Nevada and adjacent southern Cascades. The majority of collections previously associated with the epithet pinetorum represent B. pauciflora (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 401. | FNA vol. 7, p. 397. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis pygmaea | Arabis pinetorum, Arabis divaricarpa var. pinetorum, Arabis holboellii var. pinetorum, B. holboellii var. pinetorum |
Name authority | (Rollins) Al-Shehbaz: Novon 13: 388. (2003) | (Tidestrom) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 271. (2007) |
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