Boechera pygmaea |
Boechera pendulocarpa |
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Tulare County rock cress |
danglepod rockcress, drop-seed rock-cress, earless suncress, Holboell's rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases). | Perennials; short- to long-lived; sexual; caudex usually not woody. |
Stems | usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–3 dm, densely pubescent proximally, trichomes simple and stalked, 2–4-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm. |
blade narrowly oblanceolate, 1.5–4(–5) mm wide, margins entire, often ciliate along petiole, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles absent, surfaces of distalmost leaves pubescent. |
6–17, sometimes concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 2–5-flowered, unbranched. |
4–11-flowered, unbranched. |
Flowers | erect at anthesis; sepals pubescent; petals white, 3.5–5 × 0.7–1 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
arched, gently recurved, 3–8(–12) mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm; valves glabrous; ovules 8–12 per ovary; style 0.05–0.4 mm. |
pendent, not appressed to rachis, not secund, usually straight, edges parallel, (2–)2.5–3.8 cm × 1.5–2.2 mm; valves glabrous; ovules 66–92 per ovary; style 0.3–0.5 mm. |
Seeds | uniseriate, 3–5 × 2.5–4.5 mm; wing continuous, 0.8–2 mm wide distally. |
sub-biseriate, 1–1.2 × 0.7–0.9 mm; wing distal, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera pygmaea |
Boechera pendulocarpa |
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Phenology | Flowering May–Jul. | Flowering Apr–Jul. |
Habitat | Barren flats of arkosic gravel | Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows |
Elevation | 2400-3200 m (7900-10500 ft) | 1000-3300 m (3300-10800 ft) |
Distribution |
CA |
CA; ID; MT; NV; OR; UT; WY; BC; YT
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Discussion | Of conservation concern. Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993), B. pendulocarpa is easily distinguished from that species by having simple and 2–4-rayed (versus 4–8-rayed) trichomes proximally on stems, cauline leaves without auricles, fruiting pedicels gently (versus sharply) recurved, and shorter (2–)2.5–3.8 (versus 3.5–6.5) cm, non-secund fruits. The two taxa have allopatric distributions, with B. pendulocarpa found in the mountains of western North America and B. holboellii apparently confined to Greenland. Recent use of the name A. (Boechera) exilis for this taxon (e.g., G. A. Mulligan 1996; R. D. Dorn 2001; N. H. Holmgren 2005b) is based on misinterpretation of the type (M. D. Windham and I. A. Al-Shehbaz 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 401. | FNA vol. 7, p. 395. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis pygmaea | Arabis pendulocarpa, Arabis holboellii var. pendulocarpa, B. holboellii var. pendulocarpa |
Name authority | (Rollins) Al-Shehbaz: Novon 13: 388. (2003) | (A. Nelson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 77. (2006) |
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