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Graham's rockcress, Graham's suncress

Habit Perennials; usually short-lived; sexual; caudex not woody. Biennials or perennials; short-lived; apomictic; caudex present or absent.
Stems

usually 2–5 per plant, arising from margin of rosette near ground surface, (1–)2–8 dm, densely pubescent proximally, trichomes short-stalked, 3- or 4-rayed, 0.1–0.5 mm, similarly pubescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, (1.2–)2–12 dm, glabrous or pubescent proximally, trichomes simple and sessile or subsessile, 2 or 3(–6)-rayed, to 1 mm, glabrous distally.

Basal leaves

blade obovate to oblanceolate, (5–)10–45 mm wide, margins usually coarsely dentate, rarely sinuate, ciliate on petiole base, surfaces moderately pubescent, trichomes short-stalked, 3- or 4(–6)-rayed, 0.1–0.5 mm.

blade oblanceolate, 1.5–10(–20) mm wide, margins entire or dentate, ciliate proximally, trichomes (simple), to 1 mm, surfaces sparsely to densely pubescent, trichomes sessile or subsessile, 2–4(–7)-rayed, 0.2–0.6 mm.

Cauline leaves

6–25, concealing stem proximally;

blade auricles 1–6 mm, surfaces of distalmost leaves pubescent.

(10–)13–52, often concealing stem proximally;

blade auricles 1–5 mm, surfaces of distalmost leaves glabrous.

Racemes

10–40-flowered, usually unbranched.

16–88-flowered, usually unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals purplish, 3–4 × 0.7–1 mm, glabrous;

pollen ellipsoid.

divaricate at anthesis;

sepals glabrous or pubescent;

petals usually white, rarely lavender, 5.5–8 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

usually horizontal, usually straight, rarely slightly curved, 5–15 mm, pubescent, trichomes spreading, simple.

divaricate-ascending to descending, usually gently curved downward, rarely straight, 5–22(–30) mm, glabrous.

Fruits

usually horizontal, not appressed to rachis, not secund, straight, edges parallel, (1.5–)2–3.3 cm × 0.8–1 mm;

valves sparsely pubescent throughout;

ovules 12–20 per ovary;

style 0.4–0.8 mm.

divaricate-ascending to pendent, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 3.5–9 cm × 1–1.8 mm;

valves glabrous;

ovules 84–146 per ovary;

style 0.05–0.5 mm.

Seeds

uniseriate, 0.8–1.1 × 0.6–0.8 mm, not winged.

uniseriate to sub-biseriate, 1–1.8 × 0.8–1.2 mm;

wing continuous, 0.1–0.25 mm wide.

2n

= 14.

= 21.

Boechera perstellata

Boechera grahamii

Phenology Flowering Apr–May. Flowering May–Aug.
Habitat Calcareous bluffs and wooded hillsides Rocky slopes and sandy soil in prairies and open forests
Elevation 200-400 m (700-1300 ft) 100-3200 m (300-10500 ft)
Distribution
from FNA
KY; TN
from FNA
AK; CO; MI; MN; MT; ND; NY; SD; VT; WA; WI; WY; AB; BC; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Boechera perstellata is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera grahamii is an apomictic species that arose through hybridization between B. collinsii and B. stricta. Previous authors have assigned these specimens to Arabis (Boechera) divaricarpa (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison); if these taxa are treated as conspecific, the name B. grahamii has priority.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 396. FNA vol. 7, p. 382.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis perstellata, Arabis perstellata var. ampla Turritis grahamii, Arabis boivinii, Arabis bourgovii, Arabis confinis, Arabis confinis var. brachycarpa, Arabis dacotica, Arabis divaricarpa var. dacotica, Arabis divaricarpa var. hemicylindrica, Arabis divaricarpa var. stenocarpa, Arabis drummondii var. brachycarpa, Arabis holboellii var. brachycarpa, B. brachycarpa, Turritis brachycarpa
Name authority (E. L. Braun) Al-Shehbaz: Novon 13: 388. (2003) (Lehmann) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 241. (2007)
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